Latrodectus

Latrodectus is a genus of spider, in the family Theridiidae, that contains approximately 31 recognized species. The common name widow spiders is sometimes applied to members of the genus due to the supposed habit of the female of eating the male after mating, although the males of most species are not actually usually eaten after mating, and can even go on to fertilize other females. The black widow spiders are perhaps the best-known members of the genus. The injection of neurotoxic venom Latrotoxin from these species is a comparatively dangerous spider bite, resulting in the condition Latrodectism which is named for the genus. The female black widow's bite is particularly harmful to humans because of its unusually large venom glands.

Description

Not all adult female black widows exhibit the red hourglass on their abdomen—some may have a pair of red spots or have no marking at all; but any markings that are present are bright red. Adult male black widows are a quarter the size of the female, are usually gray or brown rather than black and red, while they may sometimes have an hourglass marking on their abdomen, it is usually yellow or white, not red. The bite of a male black widow is not considered dangerous to humans; it is the bite of the adult female black widow from her much-larger venom sacs that has given this spider its dangerous reputation. While there is great variation in specifics by species and by gender, any spider exhibiting a red hourglass on the abdomen and having a shiny black body is an adult female black widow.

Spiders of the genus Steatoda (also of the Theridiidae family) are often mistaken for widow spiders, and are known as "false widow spiders"; they are significantly less harmful to humans.

In common with other members of the Theridiidae family, the widow spiders construct a web, an irregular tangle of sticky silken fibers. The spider very frequently hangs upside down near the center of its web and waits for insects to blunder in and get stuck. Then, before the insect can extricate itself, the spider rushes over to bite it and wrap it in silk. If the spider perceives a threat it will quickly let itself down to the ground on a safety line of silk. As other web-weavers, these spiders have very poor eyesight and depend on vibrations reaching them through their webs to find trapped prey or warn them of larger threats. They are not aggressive; most injuries to humans are due to defensive bites delivered when a spider gets unintentionally squeezed or pinched. It is theorized that some bites may result when a spider mistakes a finger thrust into its web for its normal prey, or in cases where a female is protecting an egg sac, but ordinarily intrusion by any large creature will cause these spiders to flee.

Strength of Latrodectus silk

Silk from L. hesperus spiders is reputed to be particularly strong compared with the silk of other spiders. However, the results of a study by Blackledge, et al. do not confirm this.

The ultimate strength and other physical properties of Latrodectus hesperus (western black widow) silk were found to be similar to the properties of silk from orb-weaving spiders that had been tested in other studies. The ultimate strength for the three kinds of silk measured in the Blackledge study was about 1000 MPa. The ultimate strength reported in a previous study for Nephila edulis was 1290 MPa ± 160 MPa. The tensile strength of spider silk is comparable to that of steel wire of the same thickness; as silk is about six times less dense than steel, it is correspondingly stronger than the same weight of steel.

See the spider silk and Tensile_strength#Typical tensile strengths articles for details about the strength of spider silk in general.

Species

The southern black widows, as well as the closely related western and northern species which were previously considered the same species, have a prominent red hourglass figure on the underside of its abdomen. Many of the other widow spiders have red patterns on a glossy black or dark background, which serve as a warning. Spiders which are found in multiple regions are listed in their predominant native habitat.

Widow spiders can be found on every continent of the world except Antarctica. In North America, the black widows commonly known as Southern (Latrodectus mactans), Western (Latrodectus hesperus), and Northern (Latrodectus variolus) can be found in the United States, as can the "gray" or "brown widow spiders" (Latrodectus geometricus) and the "red widow spiders" (Latrodectus bishopi) (Preston-Malfham, 1998). The single species occurring in Australia is commonly called the Redback (Latrodectus hasselti). African species of this genus are sometimes known as button spiders.

America

The following widow spiders are indigenous to North America:

*Latrodectus bishopi, the red widow, Florida, USA

Latrodectus hesperus, the Western black widow, western Canada, United States, and Mexico.

Latrodectus mactans, the black widow spider (sometimes called the Southern black widow), warm regions of the USA.

Latrodectus variolus, the Northern black widow, from the extreme southeastern part of Canada and south to northern Florida, with frequency higher in the northern part of this range.

The following are indigenous to Central and South America.

*Latrodectus antheratus, Paraguay, Argentina

Latrodectus apicalis, Galapagos Islands

Latrodectus corallinus, Argentina

Latrodectus curacaviensis, Lesser Antilles, South America

Latrodectus diaguita, Argentina

Latrodectus mirabilis, Argentina

Latrodectus quartus, Argentina

Latrodectus variegatus, Chile and Argentina

Europe, North Africa, the Middle East, and western Asia

The following widows indigenous to the Mediterranean region, as well as in western Asia.

*Latrodectus dahli, Middle East to central Asia.

Latrodectus hystrix, Yemen, Socotra

Latrodectus lilianae, Iberian Peninsula

Latrodectus pallidus, the white widow or white steppe spider, North Africa, the Middle East, Russia, Iran, Cape Verde.

Latrodectus revivensis, Israel.

Latrodectus tredecimguttatus, the Mediterranean Black Widow or European Black Widow, Mediterranean area, central Asia, Kazakhstan. Also reported in China. Some specimens are reported as L. lugubris.

Sub-Saharan Africa and Madagascar

Latrodectus cinctus, a black button spider found in southern Africa, Cape Verde and Kuwait.

Latrodectus indistinctus, a black button spider found in South Africa and Namibia

Latrodectus karrooensis, a black button spider found in S. Africa

Latrodectus menavodi, found in Madagascar

Latrodectus obscurior, found in Cape Verde and Madagascar

Latrodectus renivulvatus, a black button spider found in Africa, Saudi Arabia and Yemen

Latrodectus rhodesiensis, a brown button spider found in Zimbabwe

South, East, and Southeast Asia

Latrodectus elegans, China, Myanmar, Japan

Latrodectus erythromelas, Sri Lanka

Latrodectus ex laos, Laos

Australia and Oceania

*Latrodectus hasselti, the redback spider, native to Australia. Also found in Southeast Asia and New Zealand, imported into both regions.

Latrodectus katipo, the red katipo, found in New Zealand

Worldwide

Latrodectus geometricus, the brown widow, grey widow, or brown button spider, Africa, USA, S. America, Australia. It is unclear where this spider originated; however it has been discovered in many warm, cosmopolitan locales.

References

Discovery Books; 2000; Insects and Spiders; St. Remy Media Inc.; New York; 35

Freeman, Scott; 2003; Biological Science; Prentice-Hall

Hillyard, Paul; 1994; the Book of Spiders; Random House, Inc.; New York; 47–50

Hillyard, Paul; 1994b; The Book of the Spider; Avon Books; New York; 22–25

Martin, Louise; 1988; Black Widow Spiders; Rourke Enterprises, Inc.; 18–20

Preston-Malfham, Ken; 1998; Spiders; Chartwell Books; Edison; New Jersey; 40.

"Arthropod," Microsoft Encarta Online Encyclopedia 2004

Abalos, J.W. (1962). The Egg-sac in the Identification of Species of Latrodectus (Black-Widow Spiders). Psyche 69:268–270. PDF

Levi, H.W. & McCrone, J.D. (1964). North American Widow Spiders of the Latrodectus curacaviensis Group (Araneae, Theridiidae. Psyche 71:12-27).PDF

External links

Tree of Life: Latrodectus

Black Widow Spider: Large format photographs and information

Description of crossing experiments between various Latrodectus species

* Category:Venomous spiders