The Kru languages belong to the Niger–Congo language family and are spoken by the Kru people from the southeast of Liberia to the east of Côte d'Ivoire. The term "Kru" is of unknown origin. According to Westermann (1952) it was used by Europeans to denote a number of tribes speaking related dialects. Marchese (1989) notes the fact that many of these peoples were recruited as “crew” by European seafarers; “the homonymy with crew is obvious, and is at least one source of the confusion among Europeans that there was a Kru/crew tribe” Andrew Dalby noted the historical importance of the Kru languages for their position at the crossroads of African-European interaction and wrote that “Kru and associated languages were among the first to be encountered by European voyagers on what was then known as the Pepper Coast, a center of the production and export of Guinea and melegueta pepper; a once staple African seaborne trade”. The Kru languages are known for some of the most complex tone systems in Africa, rivaled perhaps only by the Omotic languages.