Isoetales

Isoetales, formerly also written Isoëtales, is an order of plants in the class Isoetopsida. There are about 140-150 living species, all classified in the genus Isoetes (quillworts), with a cosmopolitan distribution but often scarce to rare. Living species are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams. Each leaf is slender and broadens downward to a swollen base up to 5 mm wide where the leaves attach in clusters to a bulb-like, underground corm characteristic of most quillwort species. This swollen base also contains male and female sporangia, protected by a thin, transparent covering (velum), which is used diagnostically to help identify quillwort species. Quillwort species are very difficult to distinguish by general appearance. The best way to identify them is by examining the megaspores under a microscope.

All quillworts and their extinct relatives are heterosporous. Some fossil species are very well known, with many stages of development and the life cycle preserved. Two of the best known are the Carboniferous Chaloneria and Cretaceous Nathorstiana.