Vestigiality describes homologous characters of organisms that have seemingly lost all or most of their original function in a species through evolution. These may take various forms such as anatomical structures, behaviors and biochemical pathways. Some of these disappear early in embryonic development, but others are retained in adulthood. Although the term serves a purpose in many gross phenotypic characteristics, vestigial has probably little applicability to DNA sequences, be they genes or non-coding sequences. Noncoding DNA has become a revealing example for vestigiality. Many types of noncoding DNA sequences have been found to have biological functions. Some sequences, however, have no known biological functions and are often referred to as "Junk DNA".
Vestigial structures are often called vestigial organs, although many of them are not actually organs. These are typically in a degenerate, atrophied, or rudimentary condition, and tend to be much more variable than similar parts. Although structures usually called "vestigial" are largely or entirely functionless, a vestigial structure may retain lesser functions or develop minor new ones. However, care must be taken not to apply the label of vestigiality to exaptation, in which a structure originally used for one purpose is modified for a new one. For example, the wings of penguins would not be vestigial, as they have been modified for a substantial new purpose (underwater locomotion), while Darwin pointed out that those of an emu would be, as they have no major function now.
Vestigial characters range on a continuum from detrimental through neutral to marginally useful. Some may be of some limited utility to an organism but still degenerate over time; the important point is not that they are without utility, but that they do not confer a significant enough advantage in terms of fitness to avoid the random force of disorder that is mutation. It is difficult, however, to say that a vestigial character is detrimental to the organism in the long term — the future is unpredictable, and that which is of no use in the present may develop into something useful in the future. Vestigiality is one of numerous lines of evidence for biological evolution.
His colleague, Jean-Baptiste Lamarck, named a number of vestigial structures in his 1809 book Philosophie Zoologique. Lamarck noted "Olivier's Spalax, which lives underground like the mole, and is apparently exposed to daylight even less than the mole, has altogether lost the use of sight: so that it shows nothing more than vestiges of this organ."
Charles Darwin was familiar with the concept of vestigial structures, though the term for them did not yet exist. He listed a number of them in The Descent of Man, including the muscles of the ear, wisdom teeth, the appendix, the tail bone, body hair, and the semilunar fold in the corner of the eye. Darwin also noted, in On the Origin of Species, that a vestigial structure could be useless for its primary function, but still retain secondary anatomical roles: "An organ serving for two purposes, may become rudimentary or utterly aborted for one, even the more important purpose, and remain perfectly efficient for the other.... [A]n organ may become rudimentary for its proper purpose, and be used for a distinct object."
In the first edition of On the Origin of Species, Darwin briefly mentioned inheritance of acquired characters under the heading "Effects of Use and Disuse", expressing little doubt that use "strengthens and enlarges certain parts, and disuse diminishes them; and that such modifications are inherited". In later editions he expanded his thoughts on this, and in the final chapter of the 6th edition concluded that species have been modified "chiefly through the natural selection of numerous successive, slight, favourable variations; aided in an important manner by the inherited effects of the use and disuse of parts".
In 1893, Robert Wiedersheim published a list of 86 human organs that were, in his words, "formerly of greater physiological significance than at present". Theorizing that they were vestiges of evolution, he called them "vestigial". Since his time, the function of some of these structures has been discovered, while other anatomical vestiges have been unearthed, making the list primarily of interest as a record of the knowledge of human anatomy at the time. Later versions of Wiedersheim's list were expanded to as many as 180 human "vestigial organs". This is why the zoologist Horatio Newman said in a written statement read into evidence in the Scopes Trial that "There are, according to Wiedersheim, no less than 180 vestigial structures in the human body, sufficient to make of a man a veritable walking museum of antiquities."
Vestigial structures are often homologous to structures that are functioning normally in other species. Therefore, vestigial structures can be considered evidence for evolution, the process by which beneficial heritable traits arise in populations over an extended period of time. The existence of vestigial traits can be attributed to changes in the environment and behavior patterns of the organism in question. As the function of the trait is no longer beneficial for survival, the likelihood that future offspring will inherit the "normal" form of it decreases. In some cases the structure becomes detrimental to the organism (for example the eyes of a mole can become infected Douglas Futuyma has stated that vestigial structures make no sense without evolution, just as spelling and usage of many modern English words can only be explained by their Latin or Old Norse antecedents.
Vestigial traits can still be considered adaptations. This is because an adaptation is often defined as a trait that has been favored by natural selection. Adaptations, therefore, need not be adaptive, as long as they were at some point.
The wings of ostriches, emus, and other flightless birds are vestigial; they are remnants of their flying ancestors' wings.
The eyes of certain cavefish and salamanders are vestigial, as they no longer allow the organism to see, and are remnants of their ancestors' functional eyes.
Boas and pythons have vestigial pelvis remnants, which are externally visible as two small pelvic spurs on each side of the cloaca. These spurs are sometimes used in copulation, but are not essential, as no colubroid snake (the vast majority of species) possesses these remnants. Furthermore, in most snakes the left lung is greatly reduced or absent. Amphisbaenians, which independently evolved limblessness, also retain vestiges of the pelvis as well as the pectoral girdle, and have lost their right lung.
Crabs have small tails tucked between their rear legs that are probably vestigial, as they are no longer in use. The working version of these tails can be found in their close crustacean relative, the lobster.
Certain species of moths (for example the Gypsy moth) have females that, although flightless, still carry small wings. These wings have no use, and are vestigial to the versions in species whose females can fly.
The fruit fly can be bred in high school experiments to produce offspring with vestigial wings, to better understand basic genetics in biology.
Other structures that still are considered vestigial include the coccyx, or tailbone (a remnant of a lost tail); the plica semilunaris on the inside corner of the eye (a remnant of the nictitating membrane); and, as pictured, muscles in the ear and other parts of the body. Other organic structures (such as the occipitofrontalis muscle) have lost their original functions (keep the head from falling) but are still useful for other purposes (facial expression).
Humans also bear some vestigial behaviors and reflexes. The formation of goose bumps in humans under stress is a vestigial reflex; its function in human ancestors was to raise the body's hair, making the ancestor appear larger and scaring off predators. Raising the hair is also used to trap an extra layer of air, keeping an animal warm. Due to the diminished amount of hair in humans, the reflex formation of goosebumps when cold is now vestigial.
Infants will instinctively grasp any object which touches the palm, in some cases strongly enough to support their own weight. It is speculated that this is because ancestral primates would have had sufficient body hair for an infant to cling to, allowing its mother to escape rapidly from danger, such as by climbing up a tree in the presence of a predator.
There are also vestigial molecular structures in humans, which are no longer in use but may indicate common ancestry with other species. One example of this is L-gulonolactone oxidase, a gene, that is functional in most other mammals, which produces an enzyme that can make vitamin C. A mutation deactivated the gene in an ancestor of the current group of primates, and it now remains in the human genome as a vestigial sequence called a pseudogene.
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Aside from the Aboriginal peoples, who according to the 2006 Canadian Census numbered 1,172,790, 3.8% of the country's total population, the majority of the populace is made up of old world immigrants and their descendents. After the initial period of French and then British colonization, different waves (or peaks) of immigration and settlement of non-aboriginal peoples took place over the course of nearly two centuries and is currently ongoing. Elements of Aboriginal, French, British and more recent immigrant customs, languages and religions have combined to form the culture of Canada and thus a Canadian identity. Canada has also been strongly influenced by that of its linguistic, geographic and economic neighbour, the United States.
The Canadian Citizenship Act of 1946 took effect on 1 January 1947. Prior to that date, Canadians were British subjects and Canada's nationality law closely mirrored that of the United Kingdom. As Canadian independence was obtained incrementally over the course of many years since the formation of the Canadian Confederation in 1867, World War I and World War II in particular gave rise to a desire amongst Canadians to have their country recognized as a fully-fledged sovereign state with a distinct citizenship. Legislation since the mid 20th century represents Canadians' commitment to multilateralism and socioeconomic development.
The British conquest of New France was proceeded by small number of Germans and Swedes who settled alongside the Scottish in Port Royal, Nova Scotia, while some Irish immigrated to the Colony of Newfoundland. In the wake of the 1775 invasion of Canada by the newly-formed Continental Army during the American Revolutionary War, approximately 60,000 United Empire Loyalist fled to British North America, a large portion of whom migrated to New Brunswick. After the War of 1812, British (included British army regulars), Scottish and Irish immigration was encouraged throughout Rupert's Land, Upper Canada and Lower Canada.
Between 1815 and 1850 some 800,000 immigrants came to the colonies of British North America, mainly from the British Isles as part of the great migration of Canada. These included some Gaelic-speaking Highland Scots displaced by the Highland Clearances to Nova Scotia. The Irish Potato Famine of the 1840s significantly increased the pace of Irish immigration to Prince Edward Island and the Province of Canada, with over 35,000 distressed individuals landing in Toronto in 1847 and 1848. Beginning in late 1850s, Chinese immigrants into the Colony of Vancouver Island and Colony of British Columbia peaked with the onset of the Fraser Canyon Gold Rush. The Chinese Immigration Act of 1885 eventually placed a head tax on all Chinese immigrants, in hopes of discouraging Chinese immigration after completion of the Canadian Pacific Railway.
The population of Canada has consistently risen, doubling approximately every 40 years, since the establishment of the Canadian Confederation in 1867. From the mid to late 19th century Canada had a policy of assisting immigrants from Europe, including an estimated 100,000 unwanted "Home Children" from Britain. Block settlement communities were established throughout western Canada between the late 19th and early 20th centuries. Some were planned and other were spontaneously created by the settlers themselves. Canada was now receiving a large amount of European immigrants predominately Italians, Germans, Scandinavians, Dutch and Ukrainians.
Legislative restrictions on immigration (such as the Continuous journey regulation and Chinese Immigration Act) that had favoured British and other European immigrants were amended in the 1960s, opening the doors to immigrants from all parts of the world. While the 1950s had still seen high levels of immigration by Europeans, by the 1970s immigrants increasingly were Chinese, Indian, Vietnamese, Jamaican and Haitian. During the late 1960s and early 1970s, Canada received many American Vietnam War draft dissenters. Throughout the late 1980s and 1990s Canada's growing Pacific trade brought with it a large influx of South Asians, that tended to settle in British Columbia.
In 2009, Canada received 252,179 immigrants - the top ten source countries were China (29,049), the Philippines (27,277), India (26,122), the United States (9,723), the United Kingdom (9,566), France (7,300), Pakistan (6,214), Iran (6,065), South Korea (5,864), and Morocco (5,222). These countries were followed closely by Algeria (4,785), United Arab Emirates (4,640), and Iraq (4,567) with Sri Lanka, Colombia, and Germany each contributing over 4,000 immigrants.
The majority of illegal immigrants come from the southern provinces of the People's Republic of China, with Asia as a whole, Eastern Europe, Caribbean, Africa and the Middle East all contributing to the illegal population. Estimates of illegal immigrants range between 35,000 and 120,000. A 2008 report by the Auditor General of Canada Sheila Fraser, stated that Canada has lost track of approximately 41,000 illegal immigrants whose visas have expired.
According to Citizenship and Immigration Canada there are three main classifications for immigrants: Family class (closely related persons of Canadian residents), Economic class (admitted on the basis of a point system that account for age, health and labour-market skills required for cost effectively inducting the immigrants into Canada's labour market) and Refugee class (those seeking protection by applying to remain in the country by way of the Canadian immigration and refugee law). In 2008, there were 65,567 immigrants in the family class, 21,860 refugees, and 149,072 economic immigrants amongst the 247,243 total immigrants to the country. Canada resettles over one in 10 of the world’s refugees and has one of the highest per-capita immigration rates in the world, driven by economic policy and family reunification, and is aiming for between 240,000 and 265,000 "new permanent residents" in 2011.
The majority of Canadian citizens live in Canada; however, there are approximately 2,800,000 Canadians abroad as of November 1, 2009. This represents about 7.5% of the total Canadian population. Of those abroad the United States, Hong Kong, the United Kingdom, Taiwan, China, and Lebanon have the largest Canadian diaspora. Canadians in United States are the greatest single expatriate community at over 1 million in 2009, representing 35.8% of all Canadians abroad. Under current Canadian law, Canada does not restrict dual citizenship but Passport Canada encourages its citizens to travel abroad on their Canadian passport, so they can access Canadian consular services .
Canadian Government policies such as; publicly-funded health care, higher and more progressive taxation, outlawing capital punishment, strong efforts to eliminate poverty, an emphasis on multiculturalism, imposing strict gun control, leniency in regard to drug use and most recently legalizing same-sex marriage - are social indicators of how Canada's political and cultural evolution differ from that of the United States. American media and entertainment are popular, if not dominant, in English Canada; conversely, many Canadian cultural products and entertainers are successful in the United States and worldwide. The Government of Canada has also influenced culture with programs, laws and institutions. It has created "crown corporations" to promote Canadian culture through media and has also tried to protect Canadian culture by setting legal minimums on Canadian content.
Canadian culture has historically been influenced by Aboriginal, French and British cultures and traditions. Most of Canada's territory was inhabited and developed later than other European colonies in the Americas, with the result that themes and symbols of pioneers, trappers, and traders were important in the early development of the Canadian identity. First Nations played a critical part in the development of European colonies in Canada, particularly for their role in assisting exploration of the continent during the North American fur trade. The British conquest of New France in the mid 1700s brought a large Francophone population under British Imperial rule, creating a need for compromise and accommodation. The new British rulers left alone much of the religious, political, and social culture of the French-speaking habitants, guaranteeing the right of the Canadiens to practice the Catholic faith and to the use of French civil law (now Quebec law) through the Quebec Act of 1774.
. Four identical sculptures are located in Buffalo City, Changchun, Sarajevo and Sydney.]]
The Constitution Act of 1867 was designed to meet the growing calls of Canadians for autonomy from British rule, while avoiding the overly-strong decentralization that contributed to the Civil War in the United States. The compromises made by the Fathers of Confederation set Canadians on a path to bilingualism, and this in turn contributed to an acceptance of diversity that later led to both multiculturalism and the recognition of Aboriginal contributions to Canadian society. Through their art and culture, First Nations, Inuit and Métis continue to exert influence on Canadian identity. however in 1917 and 1944 conscription crisis's caused a considerable rift along ethnic lines between Anglophones and Francophones. As a result of the First and Second World Wars, the Government of Canada became more assertive and less deferential to British authority. With the gradual loosening of political ties to the United Kingdom and the modernization of Canadian immigration policies, in the 20th century immigrants with African, Caribbean and Asian nationalities have added to the Canadian identity and its culture. The multiple origins immigration pattern continues today with the arrival of large numbers of immigrants from non British or French backgrounds.
Multiculturalism in Canada was adopted as the official policy of the government during the premiership of Pierre Elliot Trudeau in the 1970s and 1980s. The Canadian government has often been described as the instigator of multicultural ideology because of its public emphasis on the social importance of immigration. Multiculturalism is administered by the Department of Canadian Heritage and reflected in the law through the Canadian Multiculturalism Act and section 27 of the Canadian Charter of Rights and Freedoms. In parts of Canada, especially the major urban areas, multiculturalism itself is the cultural norm and diversity is a force that unites the community.
In a 2002 interview with the Globe and Mail, Karīm al-Hussainī the 49th Aga Khan of the Ismaili Muslims described Canada as "the most successful pluralist society on the face of our globe", citing it as "a model for the world." He explained that the experience of Canadian governance - its commitment to pluralism and its support for the rich multicultural diversity of its peoples - is something that must be shared and would be of benefit to all societies in other parts of the world. The preamble to the Canadian Charter of Rights and Freedoms references "God", and the monarch carries the title of "Defender of the Faith". However Canada has no official religion and support for religious pluralism (Freedom of religion in Canada) is an important part of Canada's political culture. With Christianity on the decline, having once been central and integral to Canadian culture and daily life; it has been recently suggested that Canada has come to enter a post-Christian period in a secular state, where the practice of religion has "moved to the margins of public life", with irreligion in Canada on the rise.
The 2001 Canadian census reported that 77.1% of Canadians identify as being Christians; of this, Catholics make up the largest group (43.6%). The largest Protestant denomination is the United Church of Canada (9.5%), followed by the Anglicans (6.8%), Baptists (2.4%), Lutherans (2%), and other Christians (4.4%). The French were first Europeans to settle in great numbers along the shores of the Saint Lawrence River, specifically Latin rite Roman Catholics, including a large number of Jesuits dedicated to converting Aboriginals; an effort that eventually proved successful. The first large Protestant communities were formed in the Maritimes after the British conquest of New France, followed by American Protestant settlers displaced by the American Revolution. The late nineteenth century saw the beginning of a large shift in Canadian immigration patterns. Large numbers of Irish and Southern Europeans immigrants were creating new Roman Catholic communities in English Canada. The settlement of the west brought significant Eastern Orthodox immigrants from Eastern Europe and Mormon and Pentecostal immigrants from the United States.
The earliest documentation of Jews in Canada are the 1754 British Army records from the French and Indian War. The Islamic, Sikhism and Buddhism communities although small, are as old as the nation itself. The 1871 Canadian Census (first "Canadian" national census) indicated thirteen Muslims among the populace, with approximately 5000 Sikh by 1908. The first Canadian mosque was constructed in Edmonton in 1938, when there were approximately 700 Muslims in Canada. Buddhism first arrived in Canada when Japanese immigrated during the late 19th century. The first Japanese Buddhist temple in Canada was built in Vancouver in 1905. The influx of immigrants in the late 20th century with Sri Lankan, Japanese, Indian and Southeast Asian customs, has contributed to the recent expansion of the Sikhism and Buddhist communities.
A multitude of languages are used by Canadians, with English and French (the official languages) being the mother tongues of 59.7% and 23.2% of the population respectively. Approximately twenty percent or over six million people in Canada list a non-official language as their mother tongue. Some of the most common first languages include: Chinese (3.1%), Italian (1.4%), German (1.2%), Spanish (1.2%), Punjabi (1.1%), Tagalog (0.9%), Tamil (0.8%), Gujarati (0.6%). Less than one percent of Canadians (just over 250,000 individuals) know how to speak an aboriginal language. About half this number (129,865) reported using an aboriginal language on a daily basis.
English and French are recognized by the Constitution of Canada as official languages. Thus all federal government laws are enacted in both English and French with government services available in both languages. In the Northwest Territories, the Official Languages Act declares that there are eleven different languages: Chipewyan, Cree, English, French, Gwich’in, Inuinnaqtun, Inuktitut, Inuvialuktun, North Slavey, South Slavey and Tłįchǫ. Canadian media offers specialty television channels, newspapers and other publications in many minority languages, that are widely accessible across the county.In Canada as elsewhere in the world of European colonization, the frontier of European exploration and settlement tended to be a linguistically diverse and fluid place, as cultures using different languages met and interacted. The need for a common means of communication between the indigenous inhabitants and new arrivals for the purposes of trade and (in some cases) intermarriage led to the development of hybrid languages. Languages like Michif, Chinook Jargon and Bungi creole tended to be highly localized and were often spoken by only a small number of individuals who were frequently capable of speaking another language.
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