Dothideomycetes is the largest and most diverse
class of
ascomycete fungi. It comprises 11
orders 90
families, 1300
genera and over 19,000 known
species. This type of development refers to the way in which the
sexual structure, bearing the sexual spores (
ascospores) forms. Dothideomycetes mostly produce flask-like structures referred to as
pseudothecia, although other shape variations do exist (e.g. see structures found in
Hysteriales). During ascolocular development pockets (locules) form first within the vegetative cells of the fungus and then all the subsequent structures form. These include the
asci which, superficially, have a thicker outer layer through which a thinner inner layer ‘bursts’, like a jack-in-a-box to release the spores. These asci are therefore referred to as bitunicate (superficially, two layers) or fissitunicate (referring to spore release). After several
DNA sequence comparisons it is now clear that another group of fungi which share these characteristics are distantly related. These are the "black yeasts" in subclass
Chaetothyriomycetidae (
Eurotiomycetes). This means that loculoascomycetes did not constitute
a natural group.
The best known members of this class are several important plant pathogens (like Phaeosphaeria nodorum and Venturia inaequalis). However, a majority of described species are either found as endophytes or saprobes growing on woody debris, decaying leaves or dung. A smaller number exist as lichens
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External links
Tree of Life Dothideomycetes
A phylogenetic re-evaluation of Dothideomycetes
Category:Ascomycota
Category:Plant pathogens and diseases