The sperm whale (Physeter macrocephalus), or cachalot, is the largest of the toothed whales and the largest toothed predator. It is the only living member of genus Physeter, and one of three extant species in the sperm whale family, along with the pygmy sperm whale and dwarf sperm whale of the genus Kogia. Mature males average at 16 metres (52 ft) in length but some may reach 20.5 metres (67 ft), with the head representing up to one-third of the animal's length. The sperm whale feeds primarily on squid. Plunging to 3 kilometres (9,800 ft) for prey, it is the deepest diving mammal. Its clicking vocalization, a form of echolocation and communication, may be as loud as 230 decibels (re 1 µPa at 1 m) underwater, making it the loudest sound produced by any animal. It has the largest brain of any animal on Earth, more than five times heavier than a human's. Sperm whales can live for more than 60 years. The sperm whale can be found anywhere in the open ocean. Females and young males live together in groups while mature males live solitary lives outside of the mating season. The females cooperate to protect and nurse their young. Females give birth every four to twenty years, and care for the calves for more than a decade. A mature sperm whale has few natural predators. Calves and weakened adults are taken by pods of orcas. From the early eighteenth century through the late 20th the species was a prime target of whalers. The head of the whale contains a liquid wax called spermaceti, from which the whale derives its name. Spermaceti was used in lubricants, oil lamps, and candles. Ambergris, a waste product from its digestive system, is still used as a fixative in perfumes. Occasionally the sperm whale's great size allowed it to defend itself effectively against whalers. The species is now protected by a whaling moratorium, and is currently listed as vulnerable by the IUCN. The sperm whale is the largest toothed whale, with adult males measuring up to 20.5 metres (67 ft) long and weighing up to 57,000 kilograms (56 long tons; 63 short tons). By contrast, the second largest toothed whale, Baird's Beaked Whale measures 12.8 metres (42 ft) and weighs up to 15 short tons (14,000 kg). The Nantucket Whaling Museum has a 5.5 metres (18 ft)-long jawbone. The museum claims that this individual was 80 feet (24 m) long; the whale that sank the Essex (one of the incidents behind Moby-Dick) was claimed to be 85 feet (26 m). A similar size is reported from a jawbone from the British Natural History Museum. A 67-foot specimen is reported from a Soviet whaling fleet near the Kurile Islands in 1950. There is disagreement on the claims of adult males approaching or exceeding 80 feet (24 m) in length. Extensive whaling may have decreased their size, as males were highly sought, primarily after World War II. Today, males do not usually exceed 18.3 metres (60 ft) in length or 51,000 kilograms (50 long tons; 56 short tons) in weight. Another view holds that exploitation by overwhaling had virtually no effect on the size of the bull sperm whales, and their size may have actually increased in current times on the basis of density dependent effects. It is among the most sexually dimorphic of all cetaceans. At birth both sexes are about the same size, but mature males are typically 30% to 50% longer and three times as massive as females. Appearance The sperm whale's unique body is unlikely to be confused with any other species. The sperm whale's distinctive shape comes from its very large, block-shaped head, which can be one-quarter to one-third of the animal's length. The S-shaped blowhole is located very close to the front of the head and shifted to the whale's left. This gives rise to a distinctive bushy, forward-angled spray. The sperm whale's flukes are triangular and very thick. Proportionally, they are larger than that of any other cetacean, and are very flexible. The whale lifts its flukes high out of the water as it begins a feeding dive. It has a series of ridges on the back's caudal third instead of a dorsal fin. The largest ridge was called the 'hump' by whalers, and can be mistaken for a dorsal fin because of its shape and size. In contrast to the smooth skin of most large whales, its back skin is usually wrinkly and has been likened to a prune by whale-watching enthusiasts. Albinos have been reported.

• published: 19 Mar 2014
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