ION Life is a digital television network carried by Ion Television affiliates, airing lifestyle programming during the day, and movies in the evening. It was launched February 19, 2007. The network airs programming pertaining to health and wellness, cooking, home decor, and travel, among other subjects.

The network is available 24 hours a day on digital subchannels (usually the third subchannel) of ION owned-and-operated stations (for example, if ION is on channel 23, ION Life would be on 23.3). ION Life programming is also carried on the ION Plus feeds available on DirecTV, some Comcast and Time Warner Cable systems, and some non-O&O stations, in place of paid programming that airs on the main ION network.

The network was going to be called iHealth prior to its rebranding as ION Television.^[1]

ION Media Networks and Comcast have reached an agreement to not only continue to carry ION Television, but also to launch two new digital networks qubo and ION Life.^[2][3]

Even though ION Life's parent network ION Television had rebranded itself on September 8, 2008, ION Life retained its present logo (which is a green version of the 2007-2008 ION Television logo) and graphics package.

Starting in February 2010, movies have been shown on ION Life, usually from 6 p.m. until 1 a.m., 7 nights a week (sometimes starting earlier or ending later depending on the length of the films). In 2012, the number of films has decreased, and more regular programming has been airing in primetime. More recently movies have been shown primarily on Monday evenings except during sweeps weeks when they are shown Monday thru Thursday evenings then the channel returns to showing Canadian imported programs.

References[link]

External links[link]

• Official website

This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed. (December 2007)

Hydrogen atom (center) contains a single proton and a single electron. Removal of the electron gives a cation (left), whereas addition of an electron gives an anion (right). The hydrogen anion, with its loosely held two-electron cloud, has a larger radius than the neutral atom, which in turn is much larger than the bare proton of the cation. Hydrogen forms the only cation that has no electrons, but even cations that (unlike hydrogen) still retain one or more electrons are still smaller than the neutral atoms or molecules from which they are derived.

An ion is an atom or molecule in which the total number of electrons is not equal to the total number of protons, giving it a net positive or negative electrical charge. The name was given by physicist Michael Faraday for the substances that allow a current to pass ("go") between electrodes in a solution, when an electric field is applied. It is the transliteration of the Greek participle ἰόν, ión, "going".

An ion consisting of a single atom is an atomic or monatomic ion; if it consists of two or more atoms, it is a molecular or polyatomic ion.

Anions and cations[link]

An anion (−) ( /ˈæn.aɪ.ən/ AN-eye-ən), from the Greek word ἄνω (ánō), meaning "up", is an ion with more electrons than protons, giving it a net negative charge (since electrons are negatively charged and protons are positively charged).

A cation (+) ( /ˈkæt.aɪ.ən/ KAT-eye-ən), from the Greek word κατά (katá), meaning "down", is an ion with fewer electrons than protons, giving it a positive charge. Since the charge on a proton is equal in magnitude to the charge on an electron, the net charge on an ion is equal to the number of protons in the ion minus the number of electrons.

General[link]

History and discovery[link]

The word ion is the Greek ιον (going), the present participle of ιεναι, ienai, "to go". This term was introduced by English physicist and chemist Michael Faraday in 1834 for the then-unknown species that goes from one electrode to the other through an aqueous medium.^[1][2] Faraday did not know the nature of these species, but he knew that since metals dissolved into and entered solution at one electrode, and new metal came forth from solution at the other electrode, that some kind of substance moved through the solution in a current, conveying matter from one place to the other.

Faraday also introduced the words anion for a negatively charged ion, and cation for a positively charged one. In Faraday's nomenclature, cations were named because they were attracted to the cathode in a galvanic device and anions were named due to their attraction to the anode.

Characteristics[link]

Ions in their gas-like state are highly reactive, and do not occur in large amounts on Earth, except in flames, lightning, electrical sparks, and other plasmas. These gas-like ions rapidly interact with ions of opposite charge to give neutral molecules or ionic salts. Ions are also produced in the liquid or solid state when salts interact with solvents (for example, water) to produce "solvated ions," which are more stable, for reasons involving a combination of energy and entropy changes as the ions move away from each other to interact with the liquid. These stabilized species are more commonly found in the environment at low temperatures. A common example is the ions present in seawater, which are derived from the dissolved salts.

All ions are charged, which means that like all charged objects they are:

• attracted to opposite electric charges (positive to negative, and vice versa),
• repelled by like charges
• when moving, travel in trajectories that are deflected by a magnetic field.

Electrons, due to their smaller mass and thus larger space-filling properties as matter waves, determine the size of atoms and molecules that possess any electrons at all. Thus, anions (negatively charged ions) are larger than the parent molecule or atom, as the excess electron(s) repel each other, and add to the physical size of the ion, because its size is determined by its electron cloud. As such, in general, cations are smaller than the corresponding parent atom or molecule due to the smaller size of its electron cloud. One particular cation (that of hydrogen) contains no electrons, and thus is very much smaller than the parent hydrogen atom.

Natural occurrences[link]

Ions are ubiquitous in nature and are responsible for diverse phenomena from the luminescence of the Sun to the existence of the Earth's ionosphere. Atoms in their ionic state may have a different color from neutral atoms, and thus light absorption by metal ions gives the color of gemstones. In both inorganic and organic chemistry (including biochemistry), the interaction of water and ions is extremely important; an example is the energy that drives breakdown of ATP. The following sections describe contexts in which ions feature prominently; these are arranged in decreasing physical length-scale, from the astronomical to the microscopic.

Astronomical[link]

The remnant of "Tycho's Supernova", a huge ball of expanding plasma. The outer shell shown in blue is X-ray emission by high-speed electrons.

A collection of non-aqueous gas-like ions, or even a gas containing a proportion of charged particles, is called a plasma. Greater than 99.9% of visible matter in the Universe may be in the form of plasmas.^[3] These include our Sun and other stars and the space between planets, as well as the space in between stars. Plasmas are often called the fourth state of matter because their properties are substantially different from those of solids, liquids, and gases. Astrophysical plasmas predominantly contain a mixture of electrons and protons (ionized hydrogen).

Related technology[link]

Ions can be non-chemically prepared using various ion sources, usually involving high voltage or temperature. These are used in a multitude of devices such as mass spectrometers, optical emission spectrometers, particle accelerators, ion implanters, and ion engines.

As reactive charged particles, they are also used in air purification by disrupting microbes, and in household items such as smoke detectors.

As signaling and metabolism in organisms are controlled by a precise ionic gradient across membranes, the disruption of this gradient contributes to cell death. This is a common mechanism exploited by natural and artificial biocides, including the ion channels gramicidin and amphotericin (a fungicide).

Inorganic dissolved ions are a component of total dissolved solids, an indicator of water quality in the world.

Chemistry[link]

Notation[link]

Denoting the charged state[link]

Equivalent notations for an iron atom (Fe) that lost two electrons.

When writing the chemical formula for an ion, its net charge is written in superscript immediately after the chemical structure for the molecule/atom. The net charge is written with the magnitude before the sign; that is, a doubly charged cation is indicated as 2+ instead of +2. However, the magnitude of the charge is omitted for singly charged molecules/atoms; for example, the sodium cation is indicated as Na⁺ and not Na¹⁺.

An alternative (and acceptable) way of showing a molecule/atom with multiple charges is by drawing out the signs multiple times; this is often seen with transition metals. Chemists sometimes circle the sign; this is merely ornamental and does not alter the chemical meaning. All three representations of Fe²⁺ shown in the figure are, thus, equivalent.

Mixed Roman numerals and charge notations for the uranyl ion. The oxidation state of the metal is shown as superscripted Roman numerals, whereas the charge of the entire complex is shown by the angle symbol together with the magnitude and sign of the net charge.

Monatomic ions are sometimes also denoted with Roman numerals; for example, the Fe²⁺ example seen above is occasionally referred to as Fe(II) or Fe^II. The Roman numeral designates the formal oxidation state of an element, whereas the superscripted numerals denotes the net charge. The two notations are, therefore, exchangeable for monatomic ions, but the Roman numerals cannot be applied to polyatomic ions. However, it is possible to mix the notations for the individual metal center with a polyatomic complex, as shown by the uranyl ion example.

Sub-classes[link]

If an ion contains unpaired electrons, it is called a radical ion. Just like uncharged radicals, radical ions are very reactive. Polyatomic ions containing oxygen, such as carbonate and sulfate, are called oxyanions. Molecular ions that contain at least one carbon to hydrogen bond are called organic ions. If the charge in an organic ion is formally centered on a carbon, it is termed a carbocation (if positively charged) or carbanion (if negatively charged).

Formation[link]

Formation of monatomic ions[link]

Monatomic ions are formed by the addition of electrons to the valence shell of the atom, which is the outer-most electron shell in an atom, or the losing of electrons from this shell. The inner shells of an atom are filled with electrons that are tightly bound to the positively charged atomic nucleus, and so do not participate in this kind of chemical interaction. The process of gaining or losing electrons from a neutral atom or molecule is called ionization.

Atoms can be ionized by bombardment with radiation, but the more usual process of ionization encountered in chemistry is the transfer of electrons between atoms or molecules. This transfer is usually driven by the attaining of stable ("closed shell") electronic configurations. Atoms will gain or lose electrons depending on which action takes the least energy.

For example, a sodium atom, Na, has a single electron in its valence shell, surrounding 2 stable, filled inner shells of 2 and 8 electrons. Since these filled shells are very stable, a sodium atom tends to lose its extra electron and attain this stable configuration, becoming a sodium cation in the process

Na → Na⁺ + e−

On the other hand, a chlorine atom, Cl, has 7 electrons in its valence shell, which is one short of the stable, filled shell with 8 electrons. Thus, a chlorine atom tends to gain an extra electron and attain a stable 8-electron configuration, becoming a chloride anion in the process:

Cl + e−
→ Cl⁻

This driving force is what causes sodium and chlorine to undergo a chemical reaction, wherein the "extra" electron is transferred from sodium to chlorine, forming sodium cations and chloride anions. Being oppositely charged, these cations and anions form ionic bonds and combine to form sodium chloride, NaCl, more commonly known as rock salt.

Na⁺ + Cl⁻ → NaCl

Formation of polyatomic and molecular ions[link]

An electrostatic potential map of the nitrate ion (NO−
3). The 3-dimensional shell represents a single arbitrary isopotential.

Polyatomic and molecular ions are often formed by the gaining or losing of elemental ions such as H⁺ in neutral molecules. For example, when ammonia, NH₃, accepts a proton, H⁺, it forms the ammonium ion, NH+
4. Ammonia and ammonium have the same number of electrons in essentially the same electronic configuration, but ammonium has an extra proton that gives it a net positive charge.

Ammonia can also lose an electron to gain a positive charge, forming the ion ·NH+
3. However, this ion is unstable, because it has an incomplete valence shell around the nitrogen atom, making it a very reactive radical ion.

Due to the instability of radical ions, polyatomic and molecular ions are usually formed by gaining or losing elemental ions such as H⁺, rather than gaining or losing electrons. This allows the molecule to preserve its stable electronic configuration while acquiring an electrical charge.

Ionization potential[link]

The energy required to detach an electron in its lowest energy state from an atom or molecule of a gas with less net electric charge is called the ionization potential, or ionization energy. The nth ionization energy of an atom is the energy required to detach its nth electron after the first n − 1 electrons have already been detached.

Each successive ionization energy is markedly greater than the last. Particularly great increases occur after any given block of atomic orbitals is exhausted of electrons. For this reason, ions tend to form in ways that leave them with full orbital blocks. For example, sodium has one valence electron in its outermost shell, so in ionized form it is commonly found with one lost electron, as Na⁺. On the other side of the periodic table, chlorine has seven valence electrons, so in ionized form it is commonly found with one gained electron, as Cl⁻. Caesium has the lowest measured ionization energy of all the elements and helium has the greatest.^[4] In general, the ionization energy of metals is much lower than the ionization energy of nonmetals, which is why, in general, metals will lose electrons to form positively charged ions and nonmetals will gain electrons to form negatively charged ions.

Ionic bonding[link]

Ionic bonding is a kind of chemical bonding that arises from the mutual attraction of oppositely charged ions. Since ions of like charge repel each other, they do not usually exist on their own. Instead, many of them may form a crystal lattice, in which ions of opposite charge are bound to each other. The resulting compound is called an ionic compound, and is said to be held together by ionic bonding. In ionic compounds there arise characteristic distances between ion neighbors from which the spatial extension and the ionic radius of individual ions may be derived.

The most common type of ionic bonding is seen in compounds of metals and nonmetals (except noble gases, which rarely form chemical compounds). Metals are characterized by having a small number of electrons in excess of a stable, closed-shell electronic configuration. As such, they have the tendency to lose these extra electrons in order to attain a stable configuration. This property is known as electropositivity. Non-metals, on the other hand, are characterized by having an electron configuration just a few electrons short of a stable configuration. As such, they have the tendency to gain more electrons in order to achieve a stable configuration. This tendency is known as electronegativity. When a highly electropositive metal is combined with a highly electronegative nonmetal, the extra electrons from the metal atoms are transferred to the electron-deficient nonmetal atoms. This reaction produces metal cations and nonmetal anions, which are attracted to each other to form a salt.

Chemical applications[link]

This section requires expansion.

Gas-like ions and solvated ions both have tremendous impact on chemical analysis and synthesis.

Mass spectrometry[link]

Catalysis[link]

Transition metal ions catalysis[link]

This section is empty. You can help by adding to it.

Templated synthesis of organic compounds[link]

This section is empty. You can help by adding to it.

Common ions[link]

See also[link]

• Air ionizer
• Anode
• Cathode
• Ion beam
• Ion source
• Ionic radius
• Auroras
• List of plasma (physics) articles

References[link]

v t e Galvanic cells Non-rechargeable: primary cells Alkaline battery Aluminium battery Bunsen cell Chromic acid cell Clark cell Daniell cell Dry cell Grove cell Leclanché cell Lithium battery Mercury battery Nickel oxyhydroxide battery Silicon–air battery Silver-oxide battery Weston cell Zamboni pile Zinc–air battery Zinc–carbon battery Zinc–chloride battery Rechargeable: secondary cells Automotive battery Lead–acid battery Lead–acid battery (gel) Lithium–air battery Lithium-ion battery Beltway battery Lithium-ion polymer battery Lithium iron phosphate battery Lithium–sulfur battery Lithium–titanate battery Nickel–cadmium battery Nickel–hydrogen battery Nickel–iron battery Nickel–lithium battery Nickel–metal hydride battery Low self-discharge NiMH battery Nickel–zinc battery Potassium-ion battery Rechargeable alkaline battery Sodium-ion battery Sodium–sulfur battery Vanadium redox battery Zinc–bromine battery Zinc–cerium battery Kinds of cells Battery Wet cell Dry cell Concentration cell Flow battery Fuel cell Trough battery Voltaic pile Parts of cells Anode Catalyst Cathode Electrolyte Half-cell Ions Salt bridge Semipermeable membrane

Life (Biota / Vitae / Eobionti)
Plants in the Rwenzori Mountains, Uganda
Scientific classification
Domains and kingdoms
Life on Earth: Non-cellular life (viruses) [note 1] Cellular life Bacteria Archaea Eukarya Protista Fungi Plantae Animalia

Life (cf. biota) is a characteristic that distinguishes objects that have signaling and self-sustaining processes from those that do not,^[1][2] either because such functions have ceased (death), or else because they lack such functions and are classified as inanimate.^[3][4] Biology is the science concerned with the study of life.

Any contiguous living system is called an organism. These animate entities undergo metabolism, maintain homeostasis, possess a capacity to grow, respond to stimuli, reproduce and, through natural selection, adapt to their environment in successive generations. More complex living organisms can communicate through various means.^[1][5] A diverse array of living organisms can be found in the biosphere of Earth, and the properties common to these organisms—plants, animals, fungi, protists, archaea, and bacteria—are a carbon- and water-based cellular form with complex organization and heritable genetic information.

Scientific evidence suggests that life began on Earth some 3.7 billion years ago. The mechanism by which life emerged is still being investigated. Since then, life has evolved into a wide variety of forms, which biologists have classified into a hierarchy of Taxa. Life can survive and thrive in a wide range of conditions. In philosophy and religion, the conception of life and its nature varies. Both offer interpretations as to how life relates to existence and consciousness, and both touch on many related issues, including life stance, purpose, conception of a god or gods, a soul or an afterlife.

Early theories[link]

Materialism[link]

Plant growth in the Hoh Rainforest

Herds of zebra and impala gathering on the Maasai Mara plain

Herds of zebra and impala gathering on the Maasai Mara plain

An aerial photo of microbial mats around the Grand Prismatic Spring of Yellowstone National Park

Some of the earliest theories of life were materialist, holding that all that exists is matter, and that all life is merely a complex form or arrangement of matter. Empedocles (430 BC) argued that every thing in the universe is made up of a combination of four eternal "elements" or "roots of all": earth, water, air, and fire. All change is explained by the arrangement and rearrangement of these four elements. The various forms of life are caused by an appropriate mixture of elements. For example, growth in plants is explained by the natural downward movement of earth and the natural upward movement of fire.^[6]

Democritus (460 BC), the disciple of Leucippus, thought that the essential characteristic of life is having a soul (psyche). In common with other ancient writers, he used the term to mean the principle of living things that causes them to function as a living thing. He thought the soul was composed of fire atoms, because of the apparent connection between life and heat, and because fire moves.^[7] He suggested that humans originally lived like animals, gradually developing communities to help one another, originating language, and developing crafts and agriculture.^[7]

In the scientific revolution of the 17th century, mechanistic ideas^{[clarification needed]} were revived by philosophers like Descartes.^{[citation needed]}

Hylomorphism[link]

Hylomorphism is the theory (originating with Aristotle (322 BC)) that all things are a combination of matter and form. Aristotle was one of the first ancient writers to approach the subject of life in a scientific way. Biology was one of his main interests, and there is extensive biological material in his extant writings. According to him, all things in the material universe have both matter and form. The form of a living thing is its soul (Greek psyche, Latin anima). There are three kinds of souls: the "vegetative soul" of plants, which causes them to grow and decay and nourish themselves, but does not cause motion and sensation; the "animal soul" which causes animals to move and feel; and the rational soul which is the source of consciousness and reasoning which (Aristotle believed) is found only in man.^[8] Each higher soul has all the attributes of the lower one. Aristotle believed that while matter can exist without form, form cannot exist without matter, and therefore the soul cannot exist without the body.^[9]

Consistent with this account is a teleological explanation of life. A teleological explanation accounts for phenomena in terms of their purpose or goal-directedness. Thus, the whiteness of the polar bear's coat is explained by its purpose of camouflage. The direction of causality is the other way round from materialistic science, which explains the consequence in terms of a prior cause. Modern biologists reject this functional view in terms of a material and causal one: biological features are to be explained not by looking forward to future optimal results, but by looking backwards to the past evolutionary history of a species, which led to the natural selection of the features in question.^{[citation needed]}

Vitalism[link]

Vitalism is the belief that the life-principle is essentially immaterial. This originated with Stahl (17th century), and held sway until the middle of the 19th century. It appealed to philosophers such as Henri Bergson, Nietzsche, Wilhelm Dilthey, anatomists like Bichat, and chemists like Liebig.^{[citation needed]}

Vitalism underpinned the idea of a fundamental separation of organic and inorganic material, and the belief that organic material can only be derived from living things. This was disproved in 1828 when Friedrich Wöhler prepared urea from inorganic materials.^[10] This so-called Wöhler synthesis is considered the starting point of modern organic chemistry. It is of historical significance because for the first time an organic compound was produced from inorganic reactants.

Later,^[when?] Helmholtz, anticipated by Mayer, demonstrated that no energy is lost in muscle movement, suggesting that there were no vital forces necessary to move a muscle. These empirical results led to the abandonment of scientific interest in vitalistic theories, although the belief lingered on in non-scientific theories such as homeopathy, which interprets diseases and sickness as caused by disturbances in a hypothetical vital force or life force.

Definitions[link]

It is a challenge for scientists and philosophers to define life in unequivocal terms.^[11][12][13] Defining life is difficult—in part—because life is a process, not a pure substance.^[14] Any definition must be sufficiently broad to encompass all life with which we are familiar, and it should be sufficiently general that, with it, scientists would not miss life that may be fundamentally different from life on Earth.^[15]

Biology[link]

Since there is no unequivocal definition of life, the current understanding is descriptive, where life is a characteristic of organisms that exhibit all or most of the following phenomena:^[14][16]

1. Homeostasis: Regulation of the internal environment to maintain a constant state; for example, electrolyte concentration or sweating to reduce temperature.
2. Organization: Being structurally composed of one or more cells, which are the basic units of life.
3. Metabolism: Transformation of energy by converting chemicals and energy into cellular components (anabolism) and decomposing organic matter (catabolism). Living things require energy to maintain internal organization (homeostasis) and to produce the other phenomena associated with life.
4. Growth: Maintenance of a higher rate of anabolism than catabolism. A growing organism increases in size in all of its parts, rather than simply accumulating matter.
5. Adaptation: The ability to change over a period of time in response to the environment. This ability is fundamental to the process of evolution and is determined by the organism's heredity as well as the composition of metabolized substances, and external factors present.
6. Response to stimuli: A response can take many forms, from the contraction of a unicellular organism to external chemicals, to complex reactions involving all the senses of multicellular organisms. A response is often expressed by motion, for example, the leaves of a plant turning toward the sun (phototropism) and by chemotaxis.
7. Reproduction: The ability to produce new individual organisms, either asexually from a single parent organism, or sexually from two parent organisms.

Alternatives[link]

To reflect the minimum phenomena required, other biological definitions of life have been proposed.^[17] Many of these proposals are based upon chemical systems. Living processes can be viewed as a delay of the spontaneous diffusion or dispersion of the internal energy of biological molecules towards more potential microstates.^[18] At a higher level, living beings are thermodynamic systems that have an organized molecular structure.^[18] Looking at it from a long-term perspective, life is matter that can reproduce itself and evolve as survival dictates.^[19][20] Hence, life is a self-sustained chemical system capable of undergoing Darwinian evolution.^[21]

Others take a systemic viewpoint that does not necessarily depend on molecular chemistry. One systemic definition of life is that living things are self-organizing and autopoietic (self-producing). Variations of this definition include Stuart Kauffman's definition as an autonomous agent or a multi-agent system capable of reproducing itself or themselves, and of completing at least one thermodynamic work cycle.^[22] Life can be modeled as a network of inferior negative feedbacks of regulatory mechanisms subordinated to a superior positive feedback formed by the potential of expansion and reproduction.^[23] From a simplified perspective, life can be said to consist of things with the capacity for metabolism and motion,^[14] or that life is self-reproduction "with variations"^[24][25] or "with an error rate below the sustainability threshold."^[25]

Biophysicists have commented on the nature and qualities of life forms—notably that they function on negative entropy.^[26][27] In more detail, according to physicists such as John Bernal, Erwin Schrödinger, Eugene Wigner, and John Avery, life is a member of the class of phenomena that are open or continuous systems able to decrease their internal entropy at the expense of substances or free energy taken in from the environment and subsequently rejected in a degraded form.^[28][29][30]

Viruses[link]

Electron micrograph of icosahedral adenovirus

Viruses are most often considered replicators rather than forms of life. They have been described as "organisms at the edge of life,"^[31] since they possess genes, evolve by natural selection,^[32] and replicate by creating multiple copies of themselves through self-assembly. However, viruses do not metabolize and they require a host cell to make new products. Virus self-assembly within host cells has implications for the study of the origin of life, as it may support the hypothesis that life could have started as self-assembling organic molecules.^[33][34]

Living systems theories[link]

Some scientists have proposed in the last few decades^[timeframe?] that a general living systems theory is required to explain the nature of life.^[35] Such a general theory, arising out of the ecological and biological sciences, attempts to map general principles for how all living systems work. Instead of examining phenomena by attempting to break things down into component parts, a general living systems theory explores phenomena in terms of dynamic patterns of the relationships of organisms with their environment.^[36]

The idea that the Earth is alive is probably as old as humankind, but the first public expression of it as a fact of science was by a Scottish scientist, James Hutton. In 1785 he stated that the Earth was a superorganism and that its proper study should be physiology. Hutton is rightly remembered as the father of geology, but his idea of a living Earth was forgotten in the intense reductionism of the 19th century.^[37] The Gaia hypothesis, originally proposed in the 1960s by scientist James Lovelock,^[38][39] explores the idea that the life on Earth functions as a single organism which defines and maintains environmental conditions necessary for its survival.^[40]

Robert Rosen (1991) built on the assumption that the explanatory powers of the mechanistic worldview cannot help understand the realm of living systems. One of several important clarifications he made was to define a system component as "a unit of organization; a part with a function, i.e., a definite relation between part and whole." From this and other starting concepts, he developed a "relational theory of systems" that attempts to explain the special properties of life. Specifically, he identified the "nonfractionability of components in an organism" as the fundamental difference between living systems and "biological machines."^[41]

A systems view of life treats environmental fluxes and biological fluxes together as a "reciprocity of influence",^[42] and a reciprocal relation with environment is arguably as important for understanding life as it is for understanding ecosystems. As Harold J. Morowitz (1992) explains it, life is a property of an ecological system rather than a single organism or species.^[43] He argues that an ecosystemic definition of life is preferable to a strictly biochemical or physical one. Robert Ulanowicz (2009) highlights mutualism as the key to understand the systemic, order-generating behavior of life and ecosystems.^[44]

Origin[link]

Evidence suggests that life on Earth has existed for about 3.7 billion years,^[45] with the oldest traces of life found in fossils dating back 3.4 billion years.^[46] All known life forms share fundamental molecular mechanisms, and based on these observations, theories on the origin of life attempt to find a mechanism explaining the formation of a primordial single cell organism from which all life originates. There are many different hypotheses regarding the path that might have been taken from simple organic molecules via pre-cellular life to protocells and metabolism. Many models fall into the "genes-first" category or the "metabolism-first" category, but a recent trend is the emergence of hybrid models that combine both categories.^[47]

There is no scientific consensus as to how life originated and all proposed theories are highly speculative. However, most accepted scientific models build in one way or another on the following hypotheses:

• The Miller-Urey experiment, and the work of Sidney Fox, suggest that conditions on the primitive Earth may have favored chemical reactions that synthesized amino acids and other organic compounds from inorganic precursors.
• Phospholipids spontaneously form lipid bilayers, the basic structure of a cell membrane.

Life as we know it today synthesizes proteins, which are polymers of amino acids using instructions encoded by cellular genes—which are polymers of deoxyribonucleic acid (DNA). Protein synthesis entails intermediary ribonucleic acid (RNA) polymers. One possibility is that genes came first and then proteins.^[48] Another possibility is that proteins came first and then genes.^[49] However, because genes are required to make proteins, and proteins are required to make genes, the problem of considering which came first is like that of the chicken or the egg. Most scientists have adopted the hypothesis that because DNA and proteins function together so intimately, it's unlikely that they arose independently.^[50] Therefore, many scientists consider the possibility, apparently first suggested by Francis Crick,^[51] that the first life was based on the DNA-protein intermediary: RNA.^[50] In fact, RNA has the DNA-like properties of information storage, replication and the catalytic properties of some proteins. Crick and others actually favored the RNA-first hypothesis^[52] even before the catalytic properties of RNA had been demonstrated by Thomas Cech.^[53]

A significant issue with the RNA-first hypothesis is that experiments designed to synthesize RNA from simple precursors have not been nearly as successful as the Miller-Urey experiments that synthesized other organic molecules from inorganic precursors. One reason for the failure to create RNA in the laboratory is that RNA precursors are very stable and do not react with each other under ambient conditions. However, the successful synthesis of certain RNA molecules under conditions hypothesized to exist prior to life on Earth has been achieved by adding alternative precursors in a specified order with the precursor phosphate present throughout the reaction.^[54] This study makes the RNA-first hypothesis more plausible to many scientists.^[55]

Recent experiments have demonstrated true Darwinian evolution of unique RNA enzymes (ribozymes) made up of two separate catalytic components that replicate each other in vitro.^[56] In describing this work from his laboratory, Gerald Joyce stated: "This is the first example, outside of biology, of evolutionary adaptation in a molecular genetic system."^[57] Such experiments make the possibility of a primordial RNA world even more attractive to many scientists.

NASA findings in 2011, based on studies with meteorites found on Earth, suggest DNA and RNA components (adenine, guanine and related organic molecules) may be formed extraterrestrially in outer space.^{[58][59][60][61]}

Conditions[link]

Cyanobacteria dramatically changed the composition of life forms on Earth by stimulating biodiversity and leading to the near-extinction of oxygen-intolerant organisms.

The diversity of life on Earth is a result of the dynamic interplay between genetic opportunity, metabolic capability, environmental challenges,^[62] and symbiosis.^[63][64][65] For most of its existence, Earth's habitable environment has been dominated by microorganisms and subjected to their metabolism and evolution. As a consequence of such microbial activities on a geologic time scale, the physical-chemical environment on Earth has been changing, thereby determining the path of evolution of subsequent life.^[62] For example, the release of molecular oxygen by cyanobacteria as a by-product of photosynthesis induced fundamental, global changes in the Earth's environment. The altered environment, in turn, posed novel evolutionary challenges to the organisms present, which ultimately resulted in the formation of our planet's major animal and plant species. Therefore this "co-evolution" between organisms and their environment is apparently an inherent feature of living systems.^[62]

All life forms require certain core chemical elements needed for biochemical functioning. These include carbon, hydrogen, nitrogen, oxygen, phosphorus, and sulfur—the elemental macronutrients for all organisms^[66]—often represented by the acronym CHNOPS. Together these make up nucleic acids, proteins and lipids, the bulk of living matter. Five of these six elements comprise the chemical components of DNA; the exception being sulfur. The latter forms a component of the important amino acid methionine. The most essential of these elements is carbon, which has the desirable attribute of forming multiple, stable covalent bonds. This allows carbon-based (organic) molecules to form an immense variety of chemical arrangements.^[67] Alternative hypothetical types of biochemistry have been proposed which eliminate one or more of these elements, swap out an element for one not on the list, or change required chiralities or other chemical properties.^{[citation needed]}

Range of tolerance[link]

The inert components of an ecosystem are the physical and chemical factors necessary for life—energy (sunlight or chemical energy), water, temperature, atmosphere, gravity, nutrients, and ultraviolet solar radiation protection.^[68] In most ecosystems the conditions vary during the day and often shift from one season to the next. To live in most ecosystems, then, organisms must be able to survive a range of conditions, called the "range of tolerance."^[69] Outside that are the "zones of physiological stress," where the survival and reproduction are possible but not optimal. Beyond these zones are the "zones of intolerance," where life for that organism is implausible. Organisms that have a wide range of tolerance are more widely distributed than organisms with a narrow range of tolerance.^[69]

Deinococcus radiodurans can resist radiation exposure.

To survive, selected microorganisms can assume forms that enable them to withstand freezing, complete desiccation, starvation, high-levels of radiation exposure, and other physical or chemical challenges. These microorganisms may survive exposure to such conditions for weeks, months, years, or even centuries.^[62] Extremophiles are microbial life forms that thrive outside the ranges where life is commonly found. They excel at exploiting uncommon sources of energy. While all organisms are composed of nearly identical molecules, evolution has enabled such microbes to cope with this wide range of physical and chemical conditions. Characterization of the structure and metabolic diversity of microbial communities in such extreme environments is ongoing.

Investigation of the tenacity and versatility of life on Earth, as well as an understanding of the molecular systems that some organisms utilize to survive such extremes, will provide a critical foundation for the search for life beyond Earth.^[62] In this regard, on April 26, 2012, scientists reported that lichen survived and showed remarkable results on the adaptation capacity of photosynthetic activity within a simulated Martian environment.^[70][71]

Organization and function[link]

Cell theory propounds the concept that cells are the basic unit of structure in every living thing. This idea was formulated by Henri Dutrochet and others during the early nineteenth century, and subsequently became widely accepted.^[72] According to this theory, all cells arise from pre-existing cells by division and the cell forms the fundamental unit of structure and function in all living organisms. The activity of an organism depends on the total activity of independent cells, with energy flow occurring within cells. Cells contain hereditary information that is carried forward as a genetic code during cell division. Cells belonging to similar species have basically the same chemical composition.

There are two primary types of cells. Prokaryotes lack a nucleus and other membrane-bound organelles, although they do they do have circular DNA and ribosomes. Bacteria and Archaea are two domains of prokaryotes. The other primary type of cells are the eukaryotes, which have distinct nuclei bound by a nuclear membrane and membrane-bound organelles, including mitochondria, chloroplasts, lysosomes, rough and smooth endoplasmic reticulum, and vacuoles. In addition, they possess organized chromosomes which store genetic material. All species of large complex organisms are eukaryotes, including animals, plants and fungi, although most species of eukaryote are protist microorganisms.^[73]

The molecular mechanisms of cell biology is based around proteins. Most of these are synthesized by the ribosomes through an enzyme-catalyzed process called protein biosynthesis. A sequence of amino acids is assembled and joined together based upon gene expression of the cell's nucleic acid. In eukaryotic cells, these proteins may then be transported and processed through the Golgi apparatus in preparation for dispatch to their destination.

Multicellular organisms may have first evolved through the formation of colonies of like cells. These cells can form a group organism through cell adhesion. The individual members of a colony are capable of surviving on their own, whereas the members of a true multi-cellular organism have developed specialties, making them dependent on the remainder of the organism for survival. Such organisms are formed from a single germ cell, which is capable of forming the various specialized cells that form the adult organism.

Classification[link]

The hierarchy of biological classification's eight major taxonomic ranks, which is an example of definition by genus and differentia. Life is divided into domains, which are subdivided into further groups. Intermediate minor rankings are not shown.

Traditionally, people have divided organisms into the classes of plants and animals, based mainly on their ability of movement. The first known attempt to classify organisms was conducted by the Greek philosopher Aristotle (384–322 BC). He classified all living organisms known at that time as either a plant or an animal. Aristotle distinguished animals with blood from animals without blood (or at least without red blood), which can be compared with the concepts of vertebrates and invertebrates respectively. He divided the blooded animals into five groups: viviparous quadrupeds (mammals), birds, oviparous quadrupeds (reptiles and amphibians), fishes and whales. The bloodless animals were divided into five groups: cephalopods, crustaceans, insects (which included the spiders, scorpions, centipedes, and what we define as insects in the present day), shelled animals (such as most molluscs and echinoderms) and "zoophytes." Though Aristotle's work in zoology was not without errors, it was the grandest biological synthesis of the time and remained the ultimate authority for many centuries after his death.^[74]

The exploration of the American continent revealed large numbers of new plants and animals that needed descriptions and classification. In the latter part of the 16th century and the beginning of the 17th, careful study of animals commenced and was gradually extended until it formed a sufficient body of knowledge to serve as an anatomical basis for classification.

In the late 1740s, Carolus Linnaeus introduced his method, still used, to formulate the scientific name of every species.^[75] Linnaeus took every effort to improve the composition and reduce the length of the many-worded names by abolishing unnecessary rhetoric, introducing new descriptive terms and defining their meaning with an unprecedented precision. By consistently using his system, Linnaeus separated nomenclature from taxonomy. This convention for naming species is referred to as binomial nomenclature.

The fungi were originally treated as plants. For a short period Linnaeus had placed them in the taxon Vermes in Animalia. He later placed them back in Plantae. Copeland classified the Fungi in his Protoctista, thus partially avoiding the problem but acknowledged their special status.^[76] The problem was eventually solved by Whittaker, when he gave them their own kingdom in his five-kingdom system. As it turned out, the fungi are more closely related to animals than to plants.^[77]

As new discoveries enabled us to study cells and microorganisms, new groups of life were revealed, and the fields of cell biology and microbiology were created. These new organisms were originally described separately in protozoa as animals and protophyta/thallophyta as plants, but were united by Haeckel in his kingdom protista, later the group of prokaryotes were split off in the kingdom Monera, eventually this kingdom would be divided in two separate groups, the Bacteria and the Archaea, leading to the six-kingdom system and eventually to the current three-domain system.^[78] The classification of eukaryotes is still controversial, with protist taxonomy especially problematic.^[79]

As microbiology, molecular biology and virology developed, non-cellular reproducing agents were discovered, such as viruses and viroids. Sometimes these entities are considered to be alive but others argue that viruses are not living organisms since they lack characteristics such as cell membrane, metabolism and do not grow or respond to their environments. Viruses can however be classed into "species" based on their biology and genetics but many aspects of such a classification remain controversial.^[80]

Since the 1960s a trend called cladistics has emerged, arranging taxa in an evolutionary or phylogenetic tree. It is unclear, should this be implemented, how the different codes will coexist.^[81]

Extraterrestrial[link]

Panspermia hypothesis

Earth is the only planet known to harbor life. The Drake equation, which predicts the number of extraterrestrial civilizations in our galaxy with which we might come in contact, has been used to discuss the probability of life elsewhere, but scientists disagree on many of the values of variables in this equation. Depending on those values, the equation may either suggest that life arises frequently or infrequently.^[91]

The region around a main sequence star that could support Earth-like life on an Earth-like planet is known as the habitable zone. The inner and outer radii of this zone vary with the luminosity of the star, as does the time interval during which the zone will survive. Stars more massive than the Sun have a larger habitable zone, but will remain on the main sequence for a shorter time interval during which life can evolve. Small red dwarf stars have the opposite problem, with a smaller habitable zone that is subject to higher levels of magnetic activity and the effects of tidal locking from close orbits. Hence, stars in the intermediate mass range such as the Sun may have a greater likelihood for Earth-like life to develop.^[92] The location of the star within a galaxy may have an impact on the likelihood of life forming. Stars in regions of a galaxy that have a greater abundance of heavier elements that can form planets, in combination with a low rate of potentially habitat-damaging supernova events, are predicted to have a higher probability of hosting planets with complex life.^[93]

Panspermia, also called exogenesis, is a hypothesis proposing that life originated elsewhere in the universe and was subsequently transferred to Earth in the form of spores perhaps via meteorites, comets or cosmic dust. In October 2011, scientists found that the cosmic dust permeating the universe contains complex organic matter ("amorphous organic solids with a mixed aromatic-aliphatic structure") that could be created naturally, and rapidly, by stars.^[94][95][96] As one of the scientists noted, "Coal and kerogen are products of life and it took a long time for them to form ... How do stars make such complicated organics under seemingly unfavorable conditions and [do] it so rapidly?"^[94] Further, the scientist suggested that these compounds may have been related to the development of life on earth and said that, "If this is the case, life on Earth may have had an easier time getting started as these organics can serve as basic ingredients for life."^[94]

Death[link]

Death is the permanent termination of all vital functions or life processes in an organism or cell.^[97][98] After death, the remains of an organism become part of the biogeochemical cycle. Organisms may be consumed by a predator or a scavenger and leftover organic material may then be further decomposed by detritivores, organisms which recycle detritus, returning it to the environment for reuse in the food chain.

One of the challenges in defining death is in distinguishing it from life. Death would seem to refer to either the moment at which life ends, or when the state that follows life begins.^[98] However, determining when death has occurred requires drawing precise conceptual boundaries between life and death. This is problematic, however, because there is little consensus over how to define life. The nature of death has for millennia been a central concern of the world's religious traditions and of philosophical inquiry. Many religions maintain faith in either a kind of afterlife or reincarnation for the soul, or resurrection of the body at a later date.

Extinction is the gradual process by which a group of taxa or species dies out, reducing biodiversity.^[99] The moment of extinction is generally considered to be the death of the last individual of that species. Because a species' potential range may be very large, determining this moment is difficult, and is usually done retrospectively after a period of apparent absence. Species become extinct when they are no longer able to survive in changing habitat or against superior competition. Over the history of the Earth, over 99% of all the species that have ever lived have gone extinct;^[100] however, mass extinctions may have accelerated evolution by providing opportunities for new groups of organisms to diversify.^[101]

Fossils are the preserved remains or traces of animals, plants, and other organisms from the remote past. The totality of fossils, both discovered and undiscovered, and their placement in fossil-containing rock formations and sedimentary layers (strata) is known as the fossil record. Such a preserved specimen is called a "fossil" if it is older than the arbitrary date of 10,000 years ago.^[102] Hence, fossils range in age from the youngest at the start of the Holocene Epoch to the oldest from the Archaean Eon, up to 3.4 billion years old.^[103][104]

See also[link]

Notes[link]

References[link]