Slug
Various species of British land slugs, including (from the top) the larger drawings: Arion ater, Kerry slug, Limax maximus and Limax flavus Arion sp., Vancouver
Scientific classification
Kingdom: Animalia Phylum: Mollusca Class: Gastropoda
Included groups
Onchidiacea Soleolifera Multiple lineages within Sigmurethra[1]

Slug is a common name that is normally applied to any gastropod mollusc that lacks a shell, that has a very reduced shell, or has a small internal shell. (This is in contrast to the common name snail, applied to gastropods that have a coiled shell large enough that the soft parts of the animal can retract fully into it.)

Slugs belong to several different lineages which also include snails that have shells. The various families of land slugs are not very closely related, despite a superficial similarity in the overall body form. The shell-less condition has arisen many times independently during the evolutionary past, and thus the category "slug" is emphatically a polyphyletic one.

As well as land slugs, there are also many marine slugs and even one freshwater slug genus (Acochlidium), but the common name "slug" is most frequently applied to air-breathing land slugs, while the marine forms are usually known as sea slugs. Land gastropods with a shell that is not quite vestigial, but is too small to retract into (like many in the family Urocyclidae), are known as semislugs.

Slugs, like all other gastropods, undergo torsion (a 180° twisting of the internal organs) during development. Internally, slug anatomy clearly shows the effects of this rotation, but externally the bodies of slugs appear rather symmetrical, except for the positioning of the pneumostome, which is on one side of the animal, normally the right hand side.

The soft, slimy bodies of slugs are prone to desiccation, so land-living slugs are confined to moist environments and must retreat to damp hiding places when the weather is dry.

Morphology and behavior[link]

Anatomy of a slug

Like other pulmonate land snails, most slugs have two pairs of 'feelers' or tentacles on their head. The upper pair is light sensing and has eyespots at the ends, while the lower pair provides the sense of smell. Both pairs are retractable, and can be regrown if lost.

On top of the slug, behind the head, is the saddle-shaped mantle, and under this are the genital opening and anus. On one side (almost always the right hand side) of the mantle is a respiratory opening, which is easy to see when open, but difficult to see when closed. This opening is known as the pneumostome. Within the mantle in some species is a very small, rather flat shell.

Like other snails, a slug moves by rhythmic waves of muscular contraction on the underside of its foot. It simultaneously secretes a layer of mucus on which it travels, which helps prevent damage to the foot tissues. Around the edge of the foot is the 'foot fringe'.

Some slug species hibernate underground during the winter in temperate climates, but in other species, the adults die in the autumn.

Vestigial shell[link]

Most slugs retain a remnant of their shell, which is usually internalized. This organ generally serves as storage for calcium salts, often in conjunction with the digestive glands.^[2]

An internal shell is present in the Limacidae^[3] and Parmacellidae.^[4]

Adult Philomycidae,^[3] Onchidiidae^[5] and Veronicellidae^[6] lack shells.

Mucus[link]

An active slug in Fremont, California

Slugs' bodies are made up mostly of water, and without a full-sized shell, their soft tissues are prone to desiccation. They must generate protective mucus to survive. Many species are most active just after rain because of the moist ground. In drier conditions, they hide in damp places such as under tree bark, fallen logs, rocks, and man-made structures, such as planters, to help retain body moisture.

Slugs produce two types of mucus: one which is thin and watery, and another which is thick and sticky. Both kinds of mucus are hygroscopic. The thin mucus spreads from the foot's centre to its edges, whereas the thick mucus spreads from front to back. Slugs also produce thick mucus which coats the whole body of the animal.

The mucus secreted by the foot contains fibres which help prevent the slug from slipping down vertical surfaces. The "slime trail" that a slug leaves behind has some secondary effects: other slugs coming across a slime trail can recognize the slime trail as produced by one of the same species, which is useful in finding a mate. Following a slime trail is also part of the hunting behavior of some carnivorous slugs.

Body mucus provides some protection against predators, as it can make the slug hard to pick up and hold by a bird's beak, for example, and the mucus itself can be distasteful.

Some species of slug secrete slime cords to lower themselves onto the ground, or to suspend a pair of slugs during copulation.

Reproduction[link]

Close-up of mating great grey slugs

Slug eggs and baby

Slugs are hermaphrodites, having both female and male reproductive organs.

Once a slug has located a mate, they encircle each other and sperm is exchanged through their protruded genitalia. A few days later the slugs lay around 30 eggs in a hole in the ground, or beneath the cover of an object such as a fallen log.

Apophallation is a commonly seen practice among many slugs. In apophallating species, the penis curls like a corkscrew and during mating often becomes entangled in the mate's genitalia. Apophallation allows the slugs to separate themselves by one or both of the slugs chewing off the other's penis. Once its penis has been removed, the slug is still able to mate using only the female parts of its reproductive system.

Ecology[link]

Arion vulgaris feeding on green leaves.

The recently discovered ghost slug hunts and eats earthworms

A. muscaria is prone to slug attack, as are many other fungi. It is unclear as to whether slugs are subject to the mushroom's drug effects.

An Arion ater slug eating an already dead earthworm.

Many slug species play an important ecosystem role by eating dead leaves, fungus, and decaying vegetable material. Other species eat parts of living plants.

Some slugs are predators and eat other slugs and snails, or earthworms.

Most carnivorous slugs on occasion also eat carrion, including dead of their own kind.

Predators[link]

Frogs, toads, snakes, hedgehogs, salamanders, eastern box turtles, rats, caecilians and also some birds and beetles are slug predators. Slugs can be a vector of transmission of parasitic nematodes that cause lungworm in various mammals, so are usually avoided by hedgehogs and other mammals when other food is available.^{[citation needed]}

Slugs, when attacked, can contract their body, making themselves harder and more compact, and combined with the slippery mucus is more difficult for many animals to grasp. The unpleasant taste of the mucus is also a deterrent.

Some slugs can self-amputate (autotomy) a portion of their tail to help the slug escape from a predator.^[7]

Human relevance[link]

The great majority of slug species are harmless to humans and to their interests, but a small number of species are serious pests of agriculture and horticulture. They can destroy foliage faster than plants can grow, thus killing even fairly large plants. They also feed on fruits and vegetables prior to harvest, making holes in the crop, which can make individual items unsuitable to sell for aesthetic reasons, and which can make the crop more vulnerable to rot and disease.

As control measures, baits are the norm in both agriculture and the garden. In recent years iron phosphate baits have emerged and are preferred over the toxic metaldehyde, especially because domestic or wild animals may be exposed to the bait. The environmentally safer iron phosphate has been shown to be at least as effective as poisonous baits.^[8] Methiocarb baits are no longer widely used.

Beneficial nematodes (Phasmarhabditis hermaphrodita) are a commercially available biological control method that are effective against a wide range of common slug species. The nematodes are applied in water and actively seek out slugs in the soil and infect them, leading to the death of the slug. This control method is suitable for use in organic growing systems.

Other slug control methods are generally ineffective, but can be somewhat useful in small gardens. These include beer traps,^[9] diatomaceous earth, crushed eggshells, coffee grounds, and copper.

It is of scientific interest that salt kills slugs by causing water to leave its body owing to osmosis ^[10] but this is not used for agricultural control as soil salinity is detrimental to crops.

In a few rare cases, humans have contracted parasite-induced meningitis from eating raw slugs.^[11]

In rural southern Italy, the garden slug Arion hortensis was used to treat gastritis, stomach ulcers or peptic ulcers by swallowing it whole and alive. Given that it is now known that most peptic ulcers are caused by Helicobacter pylori, the merit of swallowing a live slug is questionable. A clear mucus produced by the slug is also used to treat various skin conditions including dermatitis, warts, inflammations, calluses, acne and wounds.^[12]

Taxonomy[link]

Of the six orders of Pulmonata, two – the Onchidiacea and Soleolifera – solely comprise slugs. A third family, the Sigmurethra, contains various clades of snails, semi-slugs (i.e. snails whose shells are too small for them to retract fully into) and slugs.^[1]

The taxonomy of this group is in the process of being revised in the light of DNA sequencing.^[13] It appears that pulmonates are paraphyletic and basal to the opisthobranchs which are a terminal branch of the tree. The family Ellobiidae are also polyphyletic.

• Subinfraorder Orthurethra
  • Superfamily Achatinelloidea Gulick, 1873
  • Superfamily Cochlicopoidea Pilsbry, 1900
  • Superfamily Partuloidea Pilsbry, 1900
  • Superfamily Pupilloidea Turton, 1831
• Subinfraorder Sigmurethra
  • Superfamily Acavoidea Pilsbry, 1895
  • Superfamily Achatinoidea Swainson, 1840
  • Superfamily Aillyoidea Baker, 1960
  • Superfamily Arionoidea J.E. Gray in Turnton, 1840
  • Superfamily Athoracophoroidea
    • Family Athoracophoridae
  • Superfamily Orthalicoidea
    • Subfamily Bulimulinae
  • Superfamily Camaenoidea Pilsbry, 1895
  • Superfamily Clausilioidea Mörch, 1864
  • Superfamily Dyakioidea Gude & Woodward, 1921
  • Superfamily Gastrodontoidea Tryon, 1866
  • Superfamily Helicoidea Rafinesque, 1815
  • Superfamily Helixarionoidea Bourguignat, 1877
  • Superfamily Limacoidea Rafinesque, 1815
  • Superfamily Oleacinoidea H. & A. Adams, 1855
  • Superfamily Orthalicoidea Albers-Martens, 1860
  • Superfamily Plectopylidoidea Moellendorf, 1900
  • Superfamily Polygyroidea Pilsbry, 1894
  • Superfamily Punctoidea Morse, 1864
  • Superfamily Rhytidoidea Pilsbry, 1893
    • Family Rhytididae
  • Superfamily Sagdidoidera Pilsbry, 1895
  • Superfamily Staffordioidea Thiele, 1931
  • Superfamily Streptaxoidea J.E. Gray, 1806
  • Superfamily Strophocheiloidea Thiele, 1926
  • Superfamily Parmacelloidea
  • Superfamily Zonitoidea Mörch, 1864

References[link]

Further reading[link]

• Burton D. W. (January 1982). "How to be sluggish". Tuatara 25 (2): 48–63. http://www.nzetc.org/tm/scholarly/tei-Bio25Tuat02-t1-body-d2.html. 

External links[link]

Wikimedia Commons has media related to: Stylommatophora

Wikimedia Commons has media related to: Slug

Wikibooks' A Wikimanual of Gardening has more about this subject: Slugs

• Ohio State University slug fact sheet
• Pancake Slug (Veronicella sloanei) Info
• How to Get Rid of Garden Slugs- a wiki article
• Jumpings Slugs of the Pacific NW
• Land Slugs and Snails and Their Control (pub. 1959) hosted by the UNT Government Documents Department
• Slugs of Florida on the UF / IFAS Featured Creatures Web site

This biographical article needs additional citations for verification. Please help by adding reliable sources. Contentious material about living persons that is unsourced or poorly sourced must be removed immediately, especially if potentially libelous or harmful. (July 2009)

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Brother Ali
Brother Ali performing at SoundFest June 11, 2011
Background information
Birth name	Jason Newman
Born	1978 (age 33-34)
Origin	Minneapolis, Minnesota, United States
Genres	Hip hop, rap
Occupations	Rapper, Producer
Years active	1998–present
Labels	Rhymesayers Entertainment/Warner Music Group
Website	brotherali.com

Ali Newman (born Jason Newman in 1978),^[1] better known by the stage name Brother Ali, is an American hip hop artist signed to Rhymesayers Entertainment.^[2]

Personal life[link]

Brother Ali was born in Madison, Wisconsin, and spent his early childhood moving from city to city in the Midwest (mostly in Michigan). Ali's family settled in Minneapolis, Minnesota, in 1992. He attended Robbinsdale Cooper High School in New Hope, Minnesota. He has a son, Faheem, from his first marriage, and remarried in 2008. On the song "Real As Can Be" off his [EP] "Truth is Here" he says he also has a daughter on the way. According to the song "Fresh Air" on his September 2009 album Us, he goes on to say "Just got married last year/ treated so good that it ain't even fair/ already gotta boy now the baby girl's here/ Bought us a house like The Berenstein Bears." He has family in the Minneapolis area. Brother Ali takes family very seriously and sees it as a major part of life.

Ali was born with the rare genetic condition of albinism, a disorder characterized by a lack of pigment in skin, eyes, and hair. Brother Ali often makes fun of the media's constant urge to mention his condition in the first lines of their reviews or newspaper articles. He is also legally blind which is caused by his albinism.^[3]

While Brother Ali's family is white, he has often described a childhood marked by cruelty and exclusion by his white classmates as a result of his physical abnormality. He's often explained that, from an early age, he felt "most at home amongst African Americans."

When Ali's career began to blossom, he began to field questions about his race. Due to his stage name (a common reference among religious communities, particularly Muslims), many writers assumed Ali was black. When questioned on the subject, Ali declined to comment however stating in his album, "The Undisputed Truth"; "they ask me if I'm black or white, I'm neither/ race is a made up thing; I don't believe in it". He then goes on to explain how black people have played a major role in his life:

"My genes tie me to those that despised me/ Made a living killing the ones that inspired me/ I ain't just talking about singing and dancing/ I was taught life and manhood by black men/ So I'm a product of that understanding/ And a small part of me feels like I am them"

He has been practicing Islam for the past 13 years.

Career[link]

Ali's first release was a cassette-only demo tape called Rites of Passage; which Ali produced, recorded and arranged himself. The tape cemented his relationship with the local independent hip-hop label Rhymesayers Entertainment. Rites of Passage was released in 2000 and was sold almost exclusively at shows.

In 2003 Brother Ali released his first studio project, entitled Shadows on the Sun. Atmosphere producer, Ant, provided all of the instrumentals. Although the album had no major national distribution, it was heavily praised by independent hip-hop fans and publications. Ali used the album to explain his childhood and identity.

Ali's next CD was The Champion EP, released a year after Shadows on the Sun. It was presented as an extension of the full-length. Leading with a reggae inspired remix of Shadows' "Champion," the 9 song EP offered up highlights such as "Rain Water," a song about Ali losing his mother and grandfather to cancer and suicide.

In spring of 2007 Rhymesayers released The Undisputed Truth, Ali and Ant's full-length follow-up to Shadows on the Sun. "Truth" tells the story of Ali's divorce from his first wife, his struggle to gain custody of their young son, Faheem and the homelessness he faced as a single father. The first of his releases to be distributed by Warner Brothers Music, the album debuted at #69 on Billboard 200.

Brother Ali released an EP and DVD package titled "The Truth Is Here" in March 2009. The album is made up of 9 tracks including a collaboration with Rhymesayers label-mate Slug of Atmosphere.

In September 2009, Brother Ali released his follow-up to 2007's Undisputed Truth. The album was originally titled "Street Preacher" but was later changed to Us. It debuted at #56 on the Billboard 200. Ali, his DJ BK-One, and fellow Rhymesayers crew members Toki Wright and Evidence recently finished their Fresh Air Tour this fall.

Brother Ali will release his next album "Mourning in America and Dreaming in Color" on August 13th 2012.

Production[link]

Brother Ali produced the tracks "The Feeling" and "Little Girl" on Rhymesayers artist Toki Wright's debut album, "A Different Mirror."

Press[link]

Brother Ali has twice been featured in the hip-hop magazine The Source, first in August 2007, when the "Hip Hop Quotable" was from a verse in "Uncle Sam Goddamn", and second in October 2007, when he was interviewed. In May 2007, he was featured in Rolling Stone as a "New Artist to Watch". Brother Ali's personal struggle with race issues, his politics and his music are discussed in the 2011 book "From Jim Crow to Jay-Z: Race, Rap and the Performance of Masculinity".

TV appearances[link]

On August 13, 2007, Brother Ali appeared on The Late Late Show and performed his single "Uncle Sam Goddamn" from The Undisputed Truth. On October 19, 2007, Ali appeared on Late Night with Conan O'Brien and performed "Take Me Home" from The Undisputed Truth.^[4] On December 16, 2009 Ali appeared on Late Night with Jimmy Fallon and was featured with late night band, The Roots.

Inspirations[link]

Brother Ali has said many times in interviews that he finds much of his inspiration in the early 1980s rappers of the golden age of hip hop, especially KRS-One and Rakim.^[5][6]

In an interview in early 2007, Ali responded to a question about the response to his and his label's music, and the current music industry:

They just like to see something new. Like, we benefit so much right now from the fact that music sucks, and that there's so little originality. And there's so few people that are really trying to do something a little bit different so that if you're just even a little bit different...you know what I mean? People are just like "yeah," you know. As long as you're good and as long as you know how to present yourself, and, you know...but I think that people are just happy to see something a little bit different for a change, you know.^[7]

He has, however, spoken against coastal beef and mainstream vs. underground divide in hip-hop, claiming that worthwhile rap music is made by many rappers from across the spectrum, and expressing fandom and appreciation for "mainstream" artists such as Jay-Z, Rick Ross, Lil Wayne, T.I. and 50 Cent as well as respected artists who are more associated with the underground or the purist tradition, such as Mos Def, Talib Kweli, Aesop Rock, Lupe Fiasco, Jay Electronica, and his friends and frequent collaborators Murs and Joell Ortiz. Among his personal favorites are several rappers who are less acclaimed by the traditional rap purist canon, such as WC, Joe Budden, Chubb Rock and Paris.

"Uncle Sam Goddamn" Controversy[link]

Brother Ali has been under pressure from the recording industry due to lyrics from his song "Uncle Sam Goddamn", claiming creative interference from "somebody I don't wanna name, but some of you probably has their cell phones." The unnamed corporation ultimately withdrew its sponsorship of Ali, causing him to truncate parts of his 2007 tour.^[7][8] Furthermore, on the song "Second Time Around" with Benzi and Wale, Ali makes references to being kicked off a tour followed by the line "Verizon dissed me too, cuz I was too political." The song is notably critical of the United States government, with accusations that the political system is addicted to war.

Discography[link]

Studio albums[link]

EPs[link]

Singles[link]

References[link]

External links[link]

• Official Website
• Artist profile at Rhymesayers Entertainment
• Live at SXSW 2007 on 89.3 The Current
• 2009 State of Mind interview
• 2009 National Public Radio - All Things Considered interview
• "Dark Star," City Pages, April 11, 2007
• Interview on GermanRhymes.de