Moss Temporal range: Carboniferous[1] – recent PreЄ Є O S D C P T J K Pg N
"Muscinae" from Ernst Haeckel's Kunstformen der Natur, 1904
Scientific classification
Kingdom:	Plantae
Division:	Bryophyta Schimp.
Classes [2]
Takakiopsida Sphagnopsida Andreaeopsida Andreaeobryopsida Oedipodiopsida Polytrichopsida Tetraphidopsida Bryopsida

Mosses are small, soft plants that are typically 1–10 cm (0.4–4 in) tall, though some species are much larger. They commonly grow close together in clumps or mats in damp or shady locations. They do not have flowers or seeds, and their simple leaves cover the thin wiry stems. At certain times mosses produce spore capsules which may appear as beak-like capsules borne aloft on thin stalks.

There are approximately 12,000 species of moss classified in the Bryophyta.^[2] The division Bryophyta formerly included not only mosses, but also liverworts and hornworts. These other two groups of bryophytes now are often placed in their own divisions.

Physical characteristics[link]

Description[link]

Botanically, mosses are bryophytes, or non-vascular plants. They differ from 'higher' plants by not having internal water-bearing vessels or veins, and no flowers and therefore no fruits, cones or seeds. They are small (a few centimeters tall) and herbaceous (nonwoody) and absorb water and nutrients through their leaves. Mosses have stems which may be simple or branched and upright or lax, simple leaves that often have midribs, roots (rhizoids) that anchor them to their substrate, and spore-bearing capsules on long stems. They harvest sunlight to create food through photosynthesis.^[3][4] Mosses do not absorb water or nutrients from their substrate through their roots, so while mosses often grow on trees, they are never parasitic on the tree.

In addition to lacking a vascular system, mosses have a gametophyte-dominant life cycle, i.e. the plant's cells are haploid for most of its life cycle.^[5] Sporophytes (i.e. the diploid body) are short-lived and dependent on the gametophyte. This is in contrast to the pattern exhibited by most "higher" plants and by most animals. In seed plants, for example, the haploid generation is represented by the pollen and the ovule, whilst the diploid generation is the familiar flowering plant.

They can be distinguished from the similar liverworts (Marchantiophyta or Hepaticae) by their multi-cellular rhizoids. Also, in most mosses, the spore-bearing capsule enlarges and matures after its stalk elongates, while in liverworts the capsule enlarges and matures before its stalk elongates.^[4] Other differences are not universal for all mosses and all liverworts, but the presence of clearly differentiated stem with simple-shaped, ribbed leaves, without deeply lobed or segmented leaves and not arranged in three ranks, all point to the plant being a moss.

Life cycle[link]

Most kinds of plants have two sets of chromosomes in their vegetative cells and are said to be diploid, i.e. each chromosome has a partner that contains the same, or similar, genetic information. By contrast, mosses and other bryophytes have only a single set of chromosomes and so are haploid (i.e. each chromosome exists in a unique copy within the cell). There are periods in the moss life cycle when they do have a double set of paired chromosomes, but this happens only during the sporophyte stage.

Life cycle of a typical moss (Polytrichum commune)

The life of a moss starts from a haploid spore. The spore germinates to produce a protonema (pl. protonemata), which is either a mass of thread-like filaments or thalloid (flat and thallus-like). Moss protonemata typically look like a thin green felt, and may grow on damp soil, tree bark, rocks, concrete, or almost any other reasonably stable surface. This is a transitory stage in the life of a moss, but from the protonema grows the gametophore ("gamete-bearer") that is structurally differentiated into stems and leaves. A single mat of protonemata may develop several gametophore shoots, resulting in a clump of moss.

From the tips of the gametophore stems or branches develop the sex organs of the mosses. The female organs are known as archegonia (sing. archegonium) and are protected by a group of modified leaves known as the perichaetum (plural, perichaeta). The archegonia are small flask-shaped clumps of cells with an open neck (venter) down which the male sperm swim. The male organs are known as antheridia (sing. antheridium) and are enclosed by modified leaves called the perigonium (pl. perigonia). The surrounding leaves in some mosses form a splash cup, allowing the sperm contained in the cup to be splashed to neighboring stalks by falling water droplets.

Mosses can be either dioicous (compare dioecious in seed plants) or monoicous (compare monoecious). In dioicous mosses, male and female sex organs are borne on different gametophyte plants. In monoicous (also called autoicous) mosses, both are borne on the same plant. In the presence of water, sperm from the antheridia swim to the archegonia and fertilisation occurs, leading to the production of a diploid sporophyte. The sperm of mosses is biflagellate, i.e. they have two flagellae that aid in propulsion. Since the sperm must swim to the archegonium, fertilisation cannot occur without water. After fertilisation, the immature sporophyte pushes its way out of the archegonial venter. It takes about a quarter to half a year for the sporophyte to mature. The sporophyte body comprises a long stalk, called a seta, and a capsule capped by a cap called the operculum. The capsule and operculum are in turn sheathed by a haploid calyptra which is the remains of the archegonial venter. The calyptra usually falls off when the capsule is mature. Within the capsule, spore-producing cells undergo meiosis to form haploid spores, upon which the cycle can start again. The mouth of the capsule is usually ringed by a set of teeth called peristome. This may be absent in some mosses. Most mosses rely on the wind to disperse the spores. In the genus Sphagnum the spores are projected about 10 to 20 cm off the ground by compressed air contained in the capsules; the spores are accelerated to about 36,000 times the earth's gravitational acceleration g.^[6][7]

In some mosses, e.g. Ulota phyllantha, green vegetative structures called gemmae are produced on leaves or branches, which can break off and form new plants without the need to go through the cycle of fertilization. This is a means of asexual reproduction, and the genetically identical units can lead to the formation of clonal populations.

Classification[link]

Traditionally, mosses were grouped with the liverworts and hornworts in the Division Bryophyta (bryophytes), within which the mosses made up the class Musci. This definition of Bryophyta, however, is paraphyletic and now tends to be split up into three divisions. In such a system, the Division Bryophyta contains exclusively mosses.

The mosses are grouped as a single division, now named Bryophyta, and divided into eight classes:

Moss in the Allegheny National Forest, Pennsylvania, USA.

Six of the eight classes contain only one or two genera each. Polytrichopsida includes 23 genera, and Bryopsida includes the majority of moss diversity with over 95% of moss species belonging to this class.

The Sphagnopsida, the peat-mosses, comprise the two living genera Ambuchanania and Sphagnum, as well as fossil taxa. However, the genus Sphagnum is a diverse, widespread, and economically important one. These large mosses form extensive acidic bogs in peat swamps. The leaves of Sphagnum have large dead cells alternating with living photosynthetic cells. The dead cells help to store water. Aside from this character, the unique branching, thallose (flat and expanded) protonema, and explosively rupturing sporangium place it apart from other mosses.

Andreaeopsida and Andreaeobryopsida are distinguished by the biseriate (two rows of cells) rhizoids, multiseriate (many rows of cells) protonema, and sporangium that splits along longitudinal lines. Most mosses have capsules that open at the top.

Polytrichopsida have leaves with sets of parallel lamellae, flaps of chloroplast-containing cells that look like the fins on a heat sink. These carry out photosynthesis and may help to conserve moisture by partially enclosing the gas exchange surfaces. The Polytrichopsida differ from other mosses in other details of their development and anatomy too, and can also become larger than most other mosses, with e.g. Polytrichum commune forming cushions up to 40 cm (16 in) high. The tallest land moss, a member of the Polytrichidae is probably Dawsonia superba, a native to New Zealand and other parts of Australasia.

They appear to be the closest living relatives of the vascular plants.

Red moss capsules, a winter native of the Yorkshire Dales moorland.

Geological history[link]

The fossil record of moss is sparse, due to their soft-walled and fragile nature. Unambiguous moss fossils have been recovered from as early as the Permian of Antarctica and Russia, and a case is put forwards for Carboniferous mosses.^[9] It has further been claimed that tube-like fossils from the Silurian are the macerated remains of moss calyptræ.^[10]

Habitat[link]

Dense moss colonies in a cool coastal forest

A closeup of moss on a rock

Young sporophytes of the common moss Tortula muralis (wall screw-moss)

Retaining wall covered in moss

The moss garden at the Bloedel Reserve, Bainbridge Island, Washington State.

Since mosses have no vascular system to carry water through the plant, they must have a damp environment in which to live, and a surrounding of liquid water to reproduce. And since mosses are photosynthetic, they require enough sun to conduct photosynthesis.^[5] Shade tolerance varies by species, just as it does with higher plants. In most areas, mosses grow chiefly in areas of dampness and shade, such as wooded areas and at the edges of streams; but they can grow anywhere in cool damp cloudy climates, and some species are adapted to sunny, seasonally dry areas like alpine rocks or stabilized sand dunes.

Choice of substrate varies by species as well. Moss species can be classed as growing on: rocks, exposed mineral soil, disturbed soils, acid soil, calcareous soil, cliff seeps and waterfall spray areas, streamsides, shaded humusy soil, downed logs, burnt stumps, tree trunk bases, upper tree trunks, and tree branches. Moss species growing on or under trees are often specific about the species of trees they grow on, such as preferring conifers to broadleaf trees, oaks to alders, or vice versa.^[4]

Mosses are also found in cracks between paving stones in damp city streets, and on roofs. Some species adapted to disturbed, sunny areas are well adapted to urban conditions and are commonly found in cities. Examples would be Rhytidiadelphus squarrosus, a garden weed in Vancouver and Seattle areas; Bryum argenteum, the cosmopolitan sidewalk moss, and Ceratodon purpureus, red roof moss, another cosmopolitan species. A few species are wholly aquatic, such as Fontinalis antipyretica, common water moss; and others such as Sphagnum inhabit bogs, marshes and very slow-moving waterways.^[4] Such aquatic or semi-aquatic mosses can greatly exceed the normal range of lengths seen in terrestrial mosses. Individual plants 20–30 cm (8–12 in) or more long are common in Sphagnum species for example.

Wherever they occur, mosses require high levels of moisture to survive because of the lack of a vascular system, and the need for liquid water to complete fertilisation. Many mosses can survive desiccation, sometimes for months, returning to life within a few hours of rehydration.^[5]

It is generally believed that in northern latitudes, the north side of trees and rocks will generally have more luxuriant moss growth on average than other sides.^[11] This is assumed to be because the sun on the south side creates a dry environment. South of the equator the reverse would be true. However, naturalists feel that mosses grow on the damper side of trees and rocks.^[3] In some cases, such as sunny climates in temperate northern latitudes, this will be the shaded north side of the tree or rock. On steep slopes it may be the uphill side. For mosses that grow on tree branches, this is generally the upper side of the branch on horizontally growing sections or near the crotch. In cool damp cloudy climates, all sides of tree trunks and rocks may be equally damp enough for mosses. And different species of mosses have different moisture and sun requirements so will grow on different sections of the same tree or rock.

Cultivation[link]

Moss is often considered a weed in grass lawns, but is deliberately encouraged to grow under aesthetic principles exemplified by Japanese gardening. In old temple gardens, moss can carpet a forest scene. Moss is thought to add a sense of calm, age, and stillness to a garden scene. Moss is also used in bonsai to cover the soil and enhance the impression of age.^[12] Rules of cultivation are not widely established. Moss collections are quite often begun using samples transplanted from the wild in a water-retaining bag. However, specific species of moss can be extremely difficult to maintain away from their natural sites with their unique requirements of combinations of light, humidity, substrate chemistry, shelter from wind, etc.

Growing moss from spores is even less controlled. Moss spores fall in a constant rain on exposed surfaces; those surfaces which are hospitable to a certain species of moss will typically be colonised by that moss within a few years of exposure to wind and rain. Materials which are porous and moisture retentive, such as brick, wood, and certain coarse concrete mixtures are hospitable to moss. Surfaces can also be prepared with acidic substances, including buttermilk, yogurt, urine, and gently puréed mixtures of moss samples, water and ericaceous compost.

In the cool cloudy damp Pacific Northwest, moss is sometimes allowed to grow naturally as a lawn substitute, one that needs little or no mowing, fertilizing or watering. In this case, grass is considered to be the weed.^[13] Landscapers in the Seattle area sometimes collect boulders and downed logs growing mosses for installation in gardens and landscapes. Woodland gardens in many parts of the world can include a carpet of natural mosses.^[5] The Bloedel Reserve on Bainbridge Island, Washington State, is famous for its moss garden. The moss garden was created by removing shrubby underbrush and herbaceous groundcovers, thinning trees, and allowing mosses to fill in naturally.^[14]

Mosses are sometimes used in green roofs. Advantages of mosses over higher plants in green roofs include reduced weight loads, increased water absorption, no fertilizer requirements, and high drought tolerance. Since mosses do not have true roots, they require less planting medium than higher plants with extensive root systems. With proper species selection for the local climate, mosses in green roofs require no irrigation once established and are low maintenance.^[15]

Inhibiting moss growth[link]

Moss can be a troublesome weed in containerized nursery operations and greenhouses.^[16] Vigorous moss growth can inhibit seedling emergence and penetration of water and fertilizer to the plant roots.

Moss growth can be inhibited by a number of methods:

• Decreasing availability of water through drainage.
• Increasing direct sunlight.
• Increasing number and resources available for competitive plants like grasses.
• Increasing the soil pH with the application of lime.
• Heavy traffic or manually disturbing the moss bed with a rake
• Application of chemicals such as ferrous sulfate (e.g. in lawns) or bleach (e.g. on solid surfaces).
• In containerized nursery operations, coarse mineral materials such as sand, gravel, and rock chips are used as a fast-draining top dressing in plant containers to discourage moss growth.

The application of products containing ferrous sulfate or ferrous ammonium sulfate will kill moss; these ingredients are typically in commercial moss control products and fertilizers. Sulfur and Iron are essential nutrients for some competing plants like grasses. Killing moss will not prevent regrowth unless conditions favorable to their growth are changed.^[17]

Mossery[link]

A passing fad for moss-collecting in the late 19th century led to the establishment of mosseries in many British and American gardens. The mossery is typically constructed out of slatted wood, with a flat roof, open to the north side (maintaining shade). Samples of moss were installed in the cracks between wood slats. The whole mossery would then be regularly moistened to maintain growth.

Traditional uses[link]

Preindustrial societies made use of the mosses growing in their areas.

• Laplanders and other circumpolar people used mosses for bedding.^[3]
• North American tribal people used mosses for basketry, bedding, wound dressing, diapers, and menstrual fluid absorption.^[5]
• Circumpolar and alpine people used mosses as insulation in boots and mittens. Ötzi the Iceman had moss-packed boots.^[5]
• Tribes of northeastern United States and southeastern Canada used moss to fill chinks in wooden longhouses.^[5]
• Tribes of the Pacific Northwest in the US and Canada used mosses to clean salmon prior to drying, and packed wet moss into pit ovens for steaming camas bulbs. Food storage baskets and boiling baskets were also packed with mosses.^[5]

Commercial use[link]

There is a substantial market in mosses gathered from the wild. The uses for intact moss are principally in the florist trade and for home decoration. Decaying moss in the genus Sphagnum is also the major component of peat, which is "mined" for use as a fuel, as a horticultural soil additive, and in smoking malt in the production of Scotch whisky.

Sphagnum moss, generally the species cristatum and subnitens, is harvested while still growing and is dried out to be used in nurseries and horticulture as a plant growing medium. The practice of harvesting peat moss should not be confused with the harvesting of moss peat. Peat moss can be harvested on a sustainable basis and managed so that regrowth is allowed, whereas the harvesting of moss peat is generally considered to cause significant environmental damage as the peat is stripped with little or no chance of recovery.

In World War I, Sphagnum mosses were used as first-aid dressings on soldiers' wounds, as these mosses are highly absorbent and have mild antibacterial properties. Additionally, native Americans were one of the peoples to use Sphagnum for diapers and napkins, which is still done in Canada.^[18]

In rural UK, Fontinalis antipyretica was traditionally used to extinguish fires as it could be found in substantial quantities in slow-moving rivers and the moss retained large volumes of water which helped extinguish the flames. This historical use is reflected in its specific Latin/Greek name, the approximate meaning of which is "against fire".

In Finland, peat mosses have been used to make bread during famines.^[19]

In Mexico, Moss is used as a Christmas decoration.

Moss bioreactor with Physcomitrella patens

Physcomitrella patens is increasingly used in biotechnology. Prominent examples are the identification of moss genes with implications for crop improvement or human health^[20] and the safe production of complex biopharmaceuticals in the moss bioreactor, developed by Ralf Reski and his co-workers.^[21]

See also[link]

• Embryophyte
• Lichen
• Edible moss

References[link]

External links[link]

Wikimedia Commons has media related to: Bryophyta

Wikispecies has information related to: Bryophyta

Look up moss in Wiktionary, the free dictionary.

• Information, diagrams and photos
• Moss description
• Moss grower's handbook- 2.39MB, PDF file
• The British Bryological Society
• Picture Gallery of Mosses
• World of Mosses - Watercolour paintings of moss by Robert Muma

Coordinates: 56°07′02″N 3°56′23″W / 56.117222°N 3.939722°W / 56.117222; -3.939722

Stirling
Scottish Gaelic: Sruighlea
Stirling City Centre
Stirling  Stirling shown within the Stirling council area
Population	33,710 (2008)
OS grid reference	NS795935
Civil parish	Stirling
Council area	Stirling
Lieutenancy area	Stirling and Falkirk
Country	Scotland
Sovereign state	United Kingdom
Post town	STIRLING
Postcode district	FK7-FK9
Dialling code	01786
Police	Central Scotland
Fire	Central Scotland
Ambulance	Scottish
EU Parliament	Scotland
UK Parliament	Stirling
Scottish Parliament	Stirling
	Mid Scotland and Fife
List of places: UK • Scotland •

Stirling (Gaelic: Sruighlea pronounced [ˈs̪t̪ruʝlə]) is a city and former ancient burgh in Scotland, and is at the heart of the wider Stirling council area. The city is clustered around a large fortress and medieval old-town beside the River Forth. Historically it was strategically important as the "Gateway to the Highlands", with its position near the boundary between the Scottish Lowlands and Highlands, and its crossing of the Forth, the nearest to the river mouth.

Once the capital of Scotland, Stirling contains the Great Hall (restored 1999) and the Renaissance Palace (restoration completed 2011) within the Castle that rivalled any building in Europe at the time. Stirling also has its medieval parish church, The Church of the Holy Rude, where King James VI was crowned King of Scots on 29 July 1567. The Holy Rude still functions as living church with a service every Sunday.

Stirling is a centre for local government, higher education, retail, and light industry. Its population in 2008 was 33,710,^[1] for Stirling itself, the wider urban area including Bridge of Allan and Bannockburn has a population of 45,750.^[2] This makes it the smallest city in Scotland: indeed it is smaller than many of Scotland's larger towns.

One of the principal royal strongholds of the Kingdom of Scotland, Stirling was created a Royal burgh by King David I in 1130, which it remained until 1975, when the county of Stirlingshire was absorbed into Central Region. In 2002, as part of Queen Elizabeth II's Golden Jubilee, Stirling was granted city status.

History[link]

Stirling Castle (southwest aspect)

Stirling was originally a Stone Age settlement as shown by the Randolphfield standing stones and Kings Park prehistoric carvings that can still be found south of the town.^[3][4] The city has been strategically significant since at least the Roman occupation of Britain, due to its naturally defensible crag and tail hill (latterly the site of Stirling Castle), and its commanding position at the foot of the Ochil Hills on the border between the Lowlands and Highlands, at the lowest crossing point of the River Forth. It remained the river's lowest crossing until the construction of the Kincardine Bridge further downstream in the 1930s. It is supposed that Stirling is the fortress of Iuddeu or Urbs Giudi where Oswiu of Northumbria was besieged by Penda of Mercia in 655, as recorded in Bede and contemporary annals.

A ford, and later bridge, of the River Forth at Stirling brought wealth and strategic influence, as did its port. The town was chartered as a royal burgh by King David in the 12th century, with charters later reaffirmed by later monarchs (the town then referred to as Strivelyn). Major battles during the Wars of Scottish Independence took place at the Stirling Bridge in 1297 and at the nearby village of Bannockburn in 1314 involving William Wallace and Robert the Bruce respectively. There were also several Sieges of Stirling Castle in the conflict, notably in 1304. Sir Robert Felton, governor of Scarborough Castle in 1311, was slain at Stirling in 1314.

The Wallace Monument

The origin of the name Stirling is uncertain, but folk etymology suggests that it originates in either a Scots or Gaelic term meaning the place of battle, struggle or strife. Other sources suggest that it originates in a Brythonic name meaning "dwelling place of Melyn".^[5] The town has two Latin mottoes, which appeared on the earliest burgh seal of which an impression of 1296 is on record:^[6]

Hic Armis Bruti Scoti Stant Hic Cruce Tuti (The Britons stand by force of arms, The Scots are by this cross preserved from harms) and

Continet Hoc in Se Nemus et Castrum Strivilinse (The Castle and Wood of Stirling town are in the compass of this seal set down.)

Standing near the castle, the Church of the Holy Rude is one of the town's most historically important buildings. Founded in 1129 it is the second oldest building in the city after Stirling castle. It was rebuilt in the 15th century after Stirling suffered a catastrophic fire in 1405, and is reputed to be the only surviving church in the United Kingdom apart from Westminster Abbey to have held a coronation.^[7] On 29 July 1567 the infant son of Mary, Queen of Scots, was crowned James VI of Scotland here.^[7] Musket shot marks that may come from Cromwell's troops during the Wars of the Three Kingdoms are clearly visible on the tower and apse.^[7] Another important historical religious site in the area is the ruins of Cambuskenneth Abbey, the resting place of King James III of Scotland and his queen, Margaret of Denmark.^[8] During the Wars of the Three Kingdoms, the Battle of Stirling also took place in the centre of Stirling on 12 September 1648.

The tomb of James III, King of Scots at Cambuskenneth Abbey

The fortifications continued to play a strategic military role during the 18th century Jacobite Risings. In 1715, the Earl of Mar failed to take control of the castle. On 8 January 1746, the army of Bonnie Prince Charlie seized control of the town but failed to take the Castle. On their consequent retreat northwards, they blew up the church of St. Ninians where they had been storing munitions; only the tower survived and can be seen to this day.^[9]

Economically, the city's port supported overseas trade, including tea trade with India and timber trade with the Baltic. The coming of the railways in 1848 started the decline of the river trade, not least because a railway bridge downstream restricted access for shipping. By the mid 20th century the port had ceased to operate.

Famous residents have included Mary, Queen of Scots; King James VI of Scotland; Sir Henry Campbell-Bannerman; documentary film pioneer John Grierson; film music composer Muir Mathieson; animation pioneer Norman McLaren; TV presenter Kirsty Young; and footballers Billy Bremner (captain of Leeds United and Scotland) and Frank Beattie (captain of Kilmarnock).

The Barnwell brothers, Frank and Harold, worked at Grampian Motors in Causewayhead, and in 1909 they designed and flew the first powered aircraft in Scotland. Frank Barnwell went on to design aircraft including the Bristol Blenheim. A small monument to the brothers' pioneering achievement has been erected at Causewayhead roundabout.^[10]

Geography[link]

Neighbouring towns, villages and places.

	Callander	Crieff	Perth
	Loch Lomond		Alloa, Dunfermline
		Stirling    East
	Glasgow, Kirkintilloch	Cumbernauld, Denny	Falkirk, Edinburgh

Governance[link]

In terms of local government, the city of Stirling is a part of the wider Stirling Council area, which governs on matters of local administration as set out by the Local Government etc (Scotland) Act 1994. Elections to the council take place every four years. The Council is currently controlled by the Scottish National Party.^[11] The Provost of Stirling is Fergus Wood.^[12]

In terms of national government, Stirling forms part of county constituency of Stirling constituency of the House of Commons, electing one Member of Parliament (MP) to the House of Commons of the parliament of the United Kingdom by first past the post system. Anne McGuire of the Labour Party is the MP for Stirling constituency of the House of Commons.^[13]

For the purposes of the Scottish Parliament, Stirling forms part of the Stirling constituency of the Scottish Parliament constituency. The Stirling Scottish Parliament (or Holyrood) constituency created in 1999 is one of nine within the Mid Scotland and Fife electoral region. Each constituency elects one Member of the Scottish Parliament (MSP) by the first past the post system of election, and the region elects seven additional members to produce a form of proportional representation. The constituency is represented by Bruce Crawford, MSP of the Scottish National Party.^[14]

As Scotland comprises a single European Parliament Constituency, Stirling participates in electing six MEPs using the D'Hondt method of proportional representation every four years.

Geography[link]

A map of Stirling from 1945

Stirling is renowned as the Gateway to the Highlands and is generally regarded as occupying a strategic position at the point where the flatter, largely undulating Scottish Lowlands meet the rugged slopes of the Highlands along the Highland Boundary Fault.^[15][16] The starkness of this contrast is evidenced by the many hills and mountains of the lower Highlands such as Ben Vorlich and Ben Ledi which can be seen to the northwest of the city. On the other hand, the Carse of Stirling, stretching to the west and east of the city, is one of the flattest and most agriculturally productive expanses of land in the whole of Scotland.

The land surrounding Stirling has been most affected by glacial erosion and deposition. The city itself has grown up around its castle which stands atop an ancient quartz-dolerite sill, a major defensive position which was at the lowest crossing point on the River Forth. Stirling stands on the Forth at the point where the river widens and becomes tidal. To the east of the city the Ochil Hills dominate the skyline with the highest peak in the range being Ben Cleuch, although Dumyat is more noticeable from Stirling. The Ochils meet the flat carse (floodplain) of the River Forth to the east of the distinctive geographical feature of Abbey Craig, a crag and tail hill upon which stands the 220 ft (67m) high Wallace National Monument.^[17]

The climate of Stirling differs little from that of much of the rest of central Scotland. The warm Gulf Stream air current from the Atlantic Ocean is the predominant influence, with a prevailing southwesterly wind. That said, the areas round Stirling Town Centre encounter significantly less snow in Winter than many of its very close neighbours such as Denny and Dunblane. Although this could be said as being anecdotal, it is likely to be because it is at a lower level and could be said to have its own microclimate.

Areas of Stirling[link]

Top of the Town[link]