The hair cuticle is the outermost part of the hair shaft.^[1] It is a hard shingle-like layer of overlapping cells, some five to twelve deep. It is formed from dead cells which form scales that gives the hair shaft strength and do the best job of providing protection for it.^[2] The hair cuticle is the first line of defense against all forms of damage; it acts as a protective barrier for the softer inner structure including the medulla and cortex. The cuticle is responsible for much of the mechanical strength of the hair fiber. A healthy cuticle is more than just a protective layer, as the cuticle also represents the structure that controls the water content of the fiber. Much of the shine that makes healthy hair so attractive is due to the cuticle.^[3] The hair cuticle is also said to be water resistant.^[4]

Cuticles are often damaged by excessive mechanical manipulation such as brushing, using heat (like using curling irons) or chemical processing (like perms or texturizers). Everyday elements, such as the sun or wind can also cause wear and tear on your hair and damage the hair cuticles as well.^[5] Although the cuticle is the outermost layer of the hair, it does not give the hair its color because it has no melanin, which is the pigment responsible; the color of a person's hair depends on what type of melanin they have, which is found in the cortex.^[3] Also cuticle cells may be elongate, acuminate (gradually tapering to a point), ovate (egg shaped), or flattened.

References[link]

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This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed. (April 2009)

A cuticle ( /ˈkjuːtɪkəl/), or cuticola, is a term used for any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticles" are non-homologous; differing in their origin, structure, function, and chemical composition.

Human anatomy[link]

Eponychium is the anatomical term for the human nail-cuticle^{[contradictory]}

In human anatomy, cuticle (also called eponychium) refers to several structures. It refers to the layers of epidermal cells or keratinocytes that produce the horn protein keratin, and also to the superficial layer of overlapping cells covering the hair shaft (cuticula pili) that locks the hair into its follicle (See also Cuticle (hair)).

Invertebrate zoology[link]

In zoology, the invertebrate cuticle or cuticula is a multi-layered structure outside the epidermis of many invertebrates, notably roundworms^[1] and arthropods, in which it forms an exoskeleton (see arthropod exoskeleton).

The main structural components of the nematode cuticle are proteins, highly cross-linked collagens and specialised insoluble proteins known as "cuticlins", together with glycoproteins and lipids.^[2]

The main structural component of arthropod cuticle is chitin, a polysaccharide composed of N-acetylglucosamine units, together with proteins, lipids, and catecholamines The proteins and chitin are cross-linked by catecholamines such as N-acetyldopamine, contributing to their rigidity. The rigidity is a function of the types of proteins and the quantity of chitin and catecholamines. The more acidic the protein is, the softer the cuticle. It is believed that the epidermal cells and hemocytes (cells in the hemolymph) produce protein and also monitors the timing and amount of protein to be incorporated into the cuticle.^[3]

Botany[link]

In botany, plant cuticles are protective, hydrophobic, waxy coverings produced by the epidermal cells of leaves, young shoots and all other aerial plant organs. Cuticles minimize water loss and effectively reduce pathogen entry due to their waxy secretion.

The main structural components of plant cuticles are the unique polymers cutin and/or cutan, impregnated with wax.

The cuticles of plants function as permeability barriers for water and water-soluble materials. The cuticle both prevents plant surfaces from becoming wet and helps to prevent plants from drying out. Xerophytic plants such as cactus have very thick cuticles to help them survive in their arid climates. Plants that live in range of sea's spray also may have thicker cuticles that protect them from the toxic effects of salt.

Some plants, particularly those adapted to life in damp or aquatic environments, have an almost magical resistance to wetting. A well-known example is the Sacred Lotus.^[4] This spectacular adaptation is not purely the physical and chemical effect of a waxy coating however; it depends largely on the microscopic shape of the surface. When a hydrophobic surface is sculpted into microscopic, regular, elevated areas, sometimes in fractal patterns, too high and too closely spaced for the surface tension of the liquid to permit any flow into the space between the plateaus, then the area of contact between liquid and solid surfaces may be reduced to less than a tenth of what a continuous surface might permit.^[5] The effect is to reduce wetting of the surface spectacularly.^[6]

Mycology[link]

"Cuticle" is one term used for the outer layer of tissue of a mushroom's basidiocarp or "fruit body". The alternative term "pileipellis", Latin for "skin" of a "cap" (meaning "mushroom")^[7] might be technically preferable, but is perhaps too cumbersome for popular use. It is the part removed in "peeling" mushrooms. On the other hand, some morphological terminology in mycology makes finer distinctions, such as described in the article on the "pileipellis". Be that as it may, the pileipellis (or "peel") is distinct from the trama, the inner fleshy tissue of a mushroom or similar fruiting body, and also from the spore-bearing tissue layer, the hymenium.

References[link]

Wikimedia Commons has media related to: Cuticle

1. ^ About the roundworm cuticle
2. ^ Page, A.P. and Johnstone, I.L. (March 19, 2007) The cuticle, In: WormBook, ed. by Jm Kramer & D. G. Moerman. The C. elegans Research Community, WormBook, doi/10.1895/wormbook.1.138.1, [1]
3. ^ "insect physiology" The McGraw-Hill Encyclopedia of Science of Technology, Vol. 9, p. 233 2007
4. ^ Quere, D.; Surface chemistry. Fakir droplets, Nature Materials 2002, 1, 14.
5. ^ Onda T., Shibuichi S., Satoh N., Tsujii K.; "Super-Water-Repellent Fractal Surfaces"; Langmuir, 1996, 12 (9), pp 2125–2127, doi:10.1021/la950418o
6. ^ Von Baeyer, H. C., The lotus effect, The Sciences, 2000, January/February, 12
7. ^ Jaeger, Edmund C. (1959). A source-book of biological names and terms. Springfield, Ill: Thomas. ISBN 0-398-06179-3. 

Hair
Cross section of a hair
Latin	unguis
Code	TH H3.12.00.3.02001

Hair is a filamentous biomaterial, that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals, but is also found in other animals.^[1]

The human body, apart from areas of glabrous skin, is covered in follicles which produce thick terminal and fine vellus hair. Most common interest in hair is focused on hair growth, hair types and hair care, but hair is also an important biomaterial primarily composed of protein, notably keratin. In many human societies, women predominantly grow the hair on their head long while men cut theirs short.

Overview[link]

The word "hair" often refers to two distinct structures:

1. the part beneath the skin, called the hair follicle or when pulled from the skin, called the bulb. This organ is located in the dermis and maintains stem cells which not only re-grow the hair after it falls out, but also are recruited to regrow skin after a wound;^[2]
2. the shaft, which is the hard filamentous part that extends above the skin surface. A cross section of the hair shaft may be divided roughly into three zones.

Hair fibers have a structure consisting of several layers. Starting from the outside:

1. the cuticle which consists of several layers of flat, thin cells laid out overlapping one another as roof shingles,
2. the cortex, which contains the keratin bundles in cell structures that remain roughly rod-like; and in some cases,
3. the medulla, a disorganized and open area at the fiber's center.^[3]

Description[link]

Strand of human hair at 200x magnification

Each strand of hair is made up of the medulla, cortex, and cuticle.^[4] The innermost region, the medulla, is not always present and is an open, unstructured region.^[5][6] The highly structural and organized cortex, or middle layer of the hair, is the primary source of mechanical strength and water uptake. The cortex contains melanin, which colors the fiber based on the number, distribution and types of melanin granules. The shape of the follicle determines the shape of the cortex, and the shape of the fiber is related to how straight or curly the hair is. Asian hair typically has a round fiber and is quite straight. Oval and irregularly shaped fibers are generally more wavy or even curly.^[5] The cuticle is the outer covering. Its complex structure slides as the hair swells and is covered with a single molecular layer of lipid that makes the hair repel water.^[4] The diameter of human hair varies from 17 to 180 micrometers (0.00067 to 0.0071 in).^[7]

Hair growth begins inside the hair follicle. The only "living" portion of the hair is found in the follicle. The hair that is visible is the hair shaft, which exhibits no biochemical activity and is considered "dead".^[5] The base of the root is called the bulb, which contains the cells that produce the hair shaft.^[8] Other structures of the hair follicle include the oil producing sebaceous gland which lubricates the hair and the erector pili muscles, which are responsible for causing hairs to stand up. In humans, with little body hair, the effect results in goose bumps.^[9]

Natural Color[link]

All natural hair colors are the result of two types of hair pigment. Both of these pigments are melanin types, produced inside the hair follicle and packed into granules found in the fibers. Eumelanin is the dominant pigment in dark-blond, brown hair, and black hair, while pheomelanin is dominant in red hair.^[5] Blond hair is the result of having little pigmentation in the hair strand. Gray hair occurs when melanin production decreases or stops.

Human hair growth[link]

Hair grows everywhere on the external body except for mucous membranes and glabrous skin, such as that found on the palms of the hands, soles of the feet, and on the lips.

Hair follows a specific growth cycle with three distinct and concurrent phases: anagen, catagen, and telogen phases. Each has specific characteristics that determine the length of the hair. All three occur simultaneously; one strand of hair may be in the anagen phase, while another is in the telogen phase.

The body has different types of hair, including vellus hair and androgenic hair, each with its own type of cellular construction. The different construction gives the hair unique characteristics, serving specific purposes, mainly warmth and protection.

Function[link]

Many mammals have fur and other hairs that serve different functions. Hair provides thermal regulation and camouflage for many animals; for others it provides signals to other animals such as warnings, mating, or other communicative displays; and for some animals hair provides defensive functions and, rarely, even offensive protection. Hair also has a sensory function, extending the sense of touch beyond the surface of the skin. Guard hairs give warnings that may trigger a recoiling reaction.

Warmth[link]

Polar bears use their fur for warmth and while their skin is black, their transparent fur appears white and provides camouflage while hunting and serves as protection by hiding cubs in the snow.

While humans have developed clothing and other means of keeping warm, the hair found on the head serves as primary sources of heat insulation and cooling (when sweat evaporates from soaked hair) as well as protection from ultra-violet radiation exposure. The function of hair in other locations is debated. Hats and coats are still required while doing outdoor activities in cold weather to prevent frostbite and hypothermia, but the hair on the human body does help to keep the internal temperature regulated. When the body is too cold, the arrector pili muscles found attached to hair follicles stand up, causing the hair in these follicles to do the same. These hairs then form a heat-trapping layer above the epidermis. This process is formally called piloerection, derived from the Latin words 'pilus' ('hair') and 'erectio' ('stiffening'), but is more commonly known as 'having goose bumps' in humans.^[10] This is more effective in other mammals whose fur fluffs up to create air pockets between hairs that insulate the body from the cold. The opposite actions occur when the body is too warm; the arrector muscles make the hair lie flat on the skin which allows heat to leave.

Protection[link]

Human hair may not compete with the painful spines of the porcupine, but much of the hair on the human body is suited to protect it. This natural armor cannot directly protect humans from potential predators, but it does help to keep the sense organs, such as the eyes, working properly.

Touch sense[link]

Movements of hair shafts are detected by nerve receptors within the skin and by hair follicle receptors through displacement and vibration of hair shafts. Hairs can sense movements of air as well as touch by physical objects and are especially sensitive to the presence of insects. Some hairs, such as eyelashes, are especially sensitive to the presence of potentially harmful matter.^[11][12][13]

Eyebrows and eyelashes[link]

Eyelashes and eyebrows help to protect the eyes from dust, dirt, and sweat

The eyebrows provide some protection to the eyes from dirt, sweat, and rain. More importantly, they play a key role in non-verbal communication.^[14] In many other mammals they contain much longer, whisker-like hairs that act as tactile sensors.

The eyelash grows at the edges of the eyelid and protects the eye from dirt. The eyelash is to humans, camels, horses, ostriches etc., what whiskers are to cats; they are used to sense when dirt, dust, or any other potentially harmful object is too close to the eye.^[15] The eye reflexively closes as a result of this sensation.

Evolution[link]

It has been suggested that Evolution of hair be merged into this article or section. (Discuss) Proposed since March 2012.

A 2008 study by the Medical University of Vienna traced the origins of hair to the common ancestor of mammals, birds, and lizards that lived 310 million years ago. The study found chickens, lizards, and humans all possessed a similar set of genes that was involved in the production of alpha keratin. In chickens and lizards, the α-keratin produced was found in their claws, but in mammals it was used to produce hair. The scientists involved continued searching for the mechanisms that allowed mammals to use the keratins of animal claws to produce hair.^[16][17]

It was expected that dating the split of the ancestral human louse into two species, the head louse and the pubic louse, would date the loss of body hair in human ancestors. However, it turned out that the human pubic louse does not descend from the ancestral human louse, but from the gorilla louse, diverging 3.3 million years ago. This suggests that humans had lost body hair (but retained head hair) and developed thick pubic hair prior to this date, were living in or close to the forest where gorillas lived, and acquired pubic lice from butchering gorillas or sleeping in their nests.^[18][19] The evolution of the body louse from the head louse, on the other hand, places the date of clothing much later, some 100,000 years ago.^[20][21]

Human hairlessness[link]

This article or section may contain previously unpublished synthesis of published material that conveys ideas not attributable to the original sources. See the talk page for details. (November 2010)

The hairlessness of humans compared to related species may be due to loss of functionality in the pseudogene KRTHAP1 (which helps produce keratin) in the human lineage about 240,000 years ago.^[22] Mutations in the gene HR can lead to complete hair loss, though this is not typical in humans.^[23]

Humans seem hairless not because of a lack of follicles but because of the predominance of vellus fibers which are thinner, shorter, and more transparent than terminal fibers. The density of human hair follicles on the skin is about the average for an animal of equivalent size.^[24] In order to comprehend why humans are essentially hairless, it is essential to understand that mammalian body hair is not merely an aesthetic characteristic; it protects the skin from wounds, bites, heat, cold, and UV radiation.^[25] Additionally, it can be used as a communication tool and as a camouflage.^[26] To this end, it can be concluded that benefits stemming from the loss of human body hair must be great enough to outweigh the loss of these protective functions by nakedness.

Humans are the only primate species that have undergone significant hair loss and of the approximately 5000 extant species of mammal, only a handful are effectively hairless. This list includes elephants, rhinoceroses, hippopotamuses, walruses, pigs, whales and other cetaceans, and naked mole rats.^[26] Most mammals have light skin that is covered by fur, and biologists believe that early human ancestors started out this way also. Dark skin probably evolved after humans lost their body fur, because the naked skin was vulnerable to the strong UV radiation as would be experienced in Africa. Therefore, evidence of when human skin darkened has been used to date the loss of human body hair, assuming that the dark skin was needed after the fur was gone.

Dr. Alan R. Rogers, an evolutionary geneticist at the University of Utah, used mutations in the MC1R gene to estimate when human skin darkened. He said early humans may have gone through several genetic "clean sweeps" with light-skinned individuals dying off and dark-skinned individuals surviving. He estimates the last of these clean sweeps took place 1.2 million years ago.^[27] Therefore, apparent bareness in humans, likely has existed at least since that time. Later when humans began to live in more northerly latitudes, however, light skin became an advantage for the production of vitamin D and the recessive gene began to become more prevalent in those populations.

Balding, where terminal hair switches to vellus hair, usually occurs at around thirty to forty years of age. In prehistoric times, most individuals were not as likely to live past 30, with few reaching their fourth decade,^[28] and thus it wasn't likely that this trait was subject to selection.

The soft, fine hair found on many nonhuman mammals is typically called fur.

Most species evolved as the climate in Africa changed, to adjust their thermoregulation to the intense UV and sunlight at the equator, mostly by panting. Early hominids likely possessed fur similar to other large apes, but about 2.5 million years ago they developed a greater distribution of sweat glands  that enabled them to perspire over most of the body. It is not clear whether the change in body hair appearance occurred before or after the development of sweat glands. Humans have eccrine sweat glands all over their bodies.^[29] Aside from the mammary glands that produce a specialized sweat called milk, most mammals just have apocrine sweat glands on their armpits and loin. The rest of their body is covered in eccrine glands. There is a trend in primates to have increased eccrine sweat glands over the general surface of the body.^[29] It is unclear to what degree other primates sweat in response to heat, however.

The sweat glands in humans could have evolved to spread from the hands and feet as the body hair changed, or the hair change could have occurred to facilitate sweating. Horses and humans are two of the few animals capable of sweating on most of their body, yet horses are larger and still have fully developed fur. In humans, the skin hairs lie flat in hot conditions, as the erector pili muscles relax, preventing heat from being trapped by a layer of still air between the hairs, and increasing heat loss by convection.

Historically, some ideas have been advanced to explain the apparent hairlessness of humans, as compared to other species.

The thermoregulatory hypothesis (first proposed by Darwin in 1871 and later developed by Dr. Peter Wheeler, 1984, 1985) suggests that a need for decreased body hair originated as a response to climate change that began approximately 3 million years ago.^[30][31] At this time, the earth entered a period of global cooling that had a dehumidifying effect on the main early human habitats in East and Central Africa. Lush, wooded forests gave way to dry, grassland savannah; because of this, early humans were required to travel farther in search of food and water. As early humans diverged from their chimpanzee-lineage, they also became omnivorous in order to maximize calorie intake, an important distinction in a nutrient-scarce environment. Prey, however, are moving targets, and though early humans changed the traditionally ape-like appearance of the australopithecines and adapted long, strong legs to facilitate sustained running, dense, hairy coats still posed a potentially fatal risk of causing overheating during the chase.^[26] It is posited that thick hair got in the way of the sweat evaporating, so humans evolved a sparser coat of fur. Although hair provides protection against harmful UV radiation, since our hominin ancestors were bipedal, only our heads were exposed to the noonday sun.^[32] Humans kept the hair on our head which reflects harmful UV rays, but our body hair was reduced. The rise in eccrine glands occurred on the genes that determine the fate of epidermal stem cells in human embryonic development.

Another hypothesis for the thick body hair on humans proposes that Fisherian runaway sexual selection played a role (as well as in the selection of long head hair), (see types of hair and vellus hair), as well as a much larger role of testosterone in men. Sexual selection is the only theory thus far that explains the sexual dimorphism seen in the hair patterns of men and women. On average, men have more body hair than women. Males have more terminal hair, especially on the face, chest, abdomen, and back, and females have more vellus hair, which is less visible. The halting of hair development at a juvenile stage, vellus hair, would also be consistent with the neoteny evident in humans, especially in females, and thus they could have occurred at the same time.^[33] This theory, however, has significant holdings in today's cultural norms. There is no evidence that sexual selection would proceed to such a drastic extent over a million years ago when a full, lush coat of hair would most likely indicate health and would therefore be more likely to be selected for, not against, and not all human populations today have sexual dimorphism in body hair.

A final hypothesis is that human hair was reduced in response to ectoparasites.^[34][35] The “ectoparasite” explanation of modern human nakedness is based on the principle that a hairless primate would harbor fewer parasites. When our ancestors adopted group-dwelling social arrangements roughly 1.8 mya, ectoparasite loads increased dramatically. Early humans became the only one of the 193 primate species to have fleas, which can be attributed to the close living arrangements of large groups of individuals. While primate species have communal sleeping arrangements, these groups are always on the move and thus are less likely to harbor ectoparasites. Because of this, selection pressure for early humans would favor decreasing body hair because those with thick coats would have more lethal-disease-carrying ectoparasites and would thereby have lower fitness. However, early humans were not able to compensate for the loss of warmth and protection provided by body hair with clothing, and no other mammal lost body hair to reduce parasite loads.

Evolutionary variation[link]

Evolutionary biologists suggest that the genus Homo arose in East Africa approximately 2.5 million years ago.^[36] They devised new hunting techniques.^[36] The higher protein diet led to the evolution of larger body and brain sizes.^[36] Jablonski^[36] postulates that increasing body size, in conjunction with intensified hunting during the day at the equator, gave rise to a greater need to rapidly expel heat. As a result, humans evolved the ability to sweat: a process which was facilitated by the loss of body hair.^[36] A major problem with this theory, however, is that it does not explain why males are larger, hairier, and were more active in hunting than females. The female-male size differential among other closely associated primates is much greater than among humans, however, so it might have been reduced during evolution.

Other primates have sweat gland in their armpits that function as those of humans, and thus it is probable that human sweat glands evolved from a similar distribution, spreading to more areas of the body, rather than occurring through evolution of a new trait. It is not known whether the increased distribution of sweat glands occurred before, during, or after, the change in body hair, or even whether the two are related developments. Horses also sweat, and they are larger, hairier, and expend more energy running than human males, so there may not be any connection between the ability to sweat and the apparent hairlessness of humans.

Another factor in human evolution that also occurred in the prehistoric past was a preferential selection for neoteny, particularly in females. The idea that adult humans exhibit certain neotenous (juvenile) features, not evinced in the great apes, is about a century old. Louis Bolk made a long list of such traits,^[37] and Stephen Jay Gould published a short list in Ontogeny and Phylogeny.^[38] In addition, paedomorphic characteristics in women are widely acknowledged as desirable by men. For instance, vellus hair is a juvenile characteristic. However, while men develop longer, coarser, thicker, and darker terminal hair through sexual differentiation, women do not, leaving their vellus hair visible.

Texture[link]

Curly hair[link]

Jablonski^[36] asserts that it was evolutionarily advantageous for pre-humans to retain the hair on their heads in order to protect the scalp as they walked upright in the intense African (equatorial) UV light. While some might argue that, by this logic, humans should also express hairy shoulders given that these body parts would putatively be exposed to similar conditions, the protection of the head, the seat of the brain that enabled humanity to become one of the most successful species on the planet (and which also is very vulnerable at birth), was arguably a more urgent issue (axillary hair in the underarms and groin were also retained as signs of sexual maturity). During the gradual process by which Homo erectus made a transition from furry to naked skin, their hair texture putatively changed gradually from Afro-textured hair or 'kinky' (i.e. tightly coiled) to straight hair^{[citation needed]} (the condition of most mammals, including humanity's closest cousins—chimpanzees) This argument is based on the principle that curly hair impedes the passage of UV light into the body relative to straight hair (thus curly or coiled hair would be particularly advantageous for dark-skinned hominids living at the equator). It is substantiated by Iyengar's (1998) findings that UV light can enter into straight human hair roots (and thus into the body through the skin) via the hair shaft. Specifically, the results of this study suggest that this phenomenon occurs in a manner similar to the way that light passes through fiber optic tubes (which do not function as effectively when kinked or sharply curved or coiled). In this sense, during the period in which hominids (i.e. Homo Erectus) were gradually losing their straight body hair and thereby exposing the initially pale skin underneath their fur to the sun, straight hair would have been an adaptive liability as hominoids traveled farther from Africa or The Equator. Hence, straight hair may have (initially) evolved to aid the entry of UV light into the body during the transition from dark, UV-protected skin to pale-white skin.

By contrast, some believe that tightly coiled hair that grows into a typical Afro-like formation would have greatly reduced the ability of the head and brain to cool. They reason that although hair density in African peoples is much less than their European counterparts, in the intense sun the effective 'woolly hat' produced would have been a disadvantage. However, anthropologists such as Nina Jablonski make the opposite argument with regards to this hair texture. Specifically, Jablonski's assertions ^[36] suggest that the adjective "woolly" in reference to Afro-hair is a misnomer to the extent that it connotes the high heat insulation derivable from the true wool of sheep. Instead, the relatively sparse density of Afro-hair, combined with its springy coils actually results in an airy, almost sponge-like effect. This, in turn, Jablonski argues,^[36] more likely facilitates an increase in the circulation of cool air onto the scalp. Further, Afro-hair does not respond as easily to moisture and sweat as straight hair. Thus it does not stick to the neck and scalp when wet. Rather, unless totally drenched, it tends to retain its basic springy puffiness. In this sense, the trait may enhance comfort levels in intense equatorial climates compared to straight hair (which, on the other hand, tends to naturally fall over the ears and neck to a degree that provides slightly enhanced comfort levels in cold climates relative to tightly coiled hair).

Further, some interpret the ideas of Charles Darwin as suggesting that some traits, such as hair texture, were so arbitrary to human survival that the role natural selection played was trivial. Hence, they argue in favor of his suggestion that sexual selection may be responsible for such traits.^[39] However, inclinations towards deeming hair texture "adaptively trivial" may root in certain cultural value judgments more than objective logic. In this sense the possibility that hair texture may have played an adaptively significant role cannot be completely eliminated from consideration. In fact, while the sexual selection hypothesis cannot be ruled out, the asymmetrical distribution of this trait vouches for environmental influence. Specifically, if hair texture were simply the result of adaptively arbitrary human aesthetic preferences, one would expect that the global distribution of the various hair textures would be fairly random.^{[dubious – discuss]} Instead, the distribution of Afro-hair is strongly skewed toward the equator. Further, it is notable that the most pervasive expression of this hair texture can be found in sub-Saharan Africa; a region of the world that abundant genetic and paleo-anthropological evidence suggests, was the relatively recent (~200,000 year old) point of origin for modern humanity. In fact, although genetic findings (Tishkoff, 2009) suggest that sub-Saharan Africans are the most genetically diverse continental group on Earth, Afro-textured hair approaches ubiquity in this region.^{[citation needed]} This points to a strong, long-term selective pressure that, in stark contrast to most other regions of the genomes of sub-Saharan groups, left little room for genetic variation at the determining loci. Such a pattern, again, does not seem to support human sexual aesthetics as being the sole or primary cause of this distribution.

Straight hair[link]

Straight black hair

According to the recent single origin hypothesis, anatomically modern humans arose in East Africa approximately 200,000 years ago. Then, ~150,000 years later (i.e. around 50,000 years ago), sub-groups of this population began to expand our species' range to regions outside of, and (later) within, this continent (Tishkoff, 1996). For those members of this group who migrated far north (i.e. to northern Eurasia, etc.), the UV light of these regions was too weak to penetrate the highly pigmented skin of the initially (relatively) dark-skinned migrants so as to provide enough vitamin D for healthy bone development.^[36] Malformed bones in the pelvic area were especially deadly for women because they interfered with the successful delivery of babies, leading to the death of both the mother and the infant during labor. Hence, those with less pigmented skin survived and had children at higher rates because their skin allowed more UV light for the production of vitamin D.^[36] Thus, the skin of those in the group that left the African continent and went far north gradually developed adaptations for relatively greater translucence compared to equatorial hues. This enabled the passage of more UV light into the body at high latitudes, facilitating the natural human body-process of manufacturing vitamin D (which is essential for bone development) in response to said light.^[36]

In this sense, the evidence with regard to the evolution of straight hair texture seems to support Jablonski's suggestions ^[36] that the need for vitamin D triggered the transition from dark to pale, translucent skin among modern humans. Specifically, the distribution of this trait suggests that this need may have (initially) grown so intense at certain (early) points that those among said (initially more deeply pigmented skinned) Northern-migrants with mutations for straighter hair survived and had children at (somewhat) higher rates. This early change in texture was likely subsequently followed by the accumulation of adaptively advantageous genetic changes that led to the above-mentioned skin-translucence. This argument is made based on the principle that straight fibers better facilitate the passage of UV light into the body relative to curly hair. It is substantiated by Iyengar's (1998) findings that UV light can pass through straight human hair roots in a manner similar to the way that light passes through fiber optic tubes (Iyengar, 1998).

Man with straight hair

Nonetheless, some argue against this stance because straighter hair ends tend to point downward while fiber optics requires that light be transmitted at a high angle to the normal of the inner reflective surface. In light of this, they suggest that only light reflected from the ground could successfully enter the hair follicle and be transmitted down the shaft. Even this process, they argue, is hindered by the curvature at the base of the hair. Therefore, coupled with the amount of skin covered by long head hair, these factors seem to militate against the adaptive usefulness of straight hair at northern latitudes. They further argue that UV light also is poorly reflected from soil and dull surfaces. These ideas can be countered by the fact that during the winter, the time of year in which UV light is most scarce at northern latitudes, the ground is often covered with white snow. Given that white is the most effective color in terms of facilitating the reflection of ground light, the hypothesis that straight hair could have been adaptively favorable, cannot be fully discounted in this regard. In addition, as mentioned in the previous section, straight hair also may have contributed to enhanced comfort levels in the north. This is evident in the extent to which, relative to curly hair, it tends to provide a layer of protection for ears and necks against the cold.

The latter hypothesis seems the more plausible evolution determinant as the surface area of the head is minute compared to the remainder of the body, thus the energy required in producing long hair for the express purpose of "optical" amplification of UV light reflected from the snow seems counterproductive (however, it's very likely that the trait was sustained due to a nuanced combination of multiple influences, given that human hunting-skills and ingenuity were such by 50,000 years ago that said benefits in terms of 'comfort' could have alternatively been derived from constructing head and ear warmers of fur from prey, etc.). Scientists point to the fact that straight hair found in many ethnic groups is denser as well and has a greater ability to "show" as it does not coil, hence providing more warmth as the likely deterministic factor for the evolution of straight long hair. Some scientists argue that since the head and appendages are the greatest areas for heat loss from the body, the ability to grow long hair on the crown of the head as well as the face provides a distinct advantage in a cold climate. Since the main sensory organs are anatomically located on the head, long hair provides the necessary warmth and protection in a cold climate that allows the use of these organs by exposing them to the elements to "sense", in for example a hunt, yet still providing necessary warmth and protection to sustain prolonged exposure. It may be argued, therefore, that the ability to grow long, straight, densely packed hair provides a distinct evolutionary advantage in cold climate; however, it would be a distinct disadvantage in a hot climate, when compared to loosely packed, spongy, closely cropped hair.

The EDAR Locus[link]

A group of studies have recently shown that genetic patterns at the EDAR locus, a region of the modern human genome that contributes to hair texture variation among most individuals of East Asian descent, support the hypothesis that (East Asian) straight hair likely developed in this branch of the modern human lineage subsequent to the original expression of tightly coiled natural afro-hair.^[40][41][42] Specifically, the relevant findings indicate that the EDAR mutation coding for the predominant East Asian 'coarse' or thick, straight hair texture arose within the past ~65,000 years, which is a time frame that covers from the earliest of the 'Out of Africa' migrations up to now.

Removal practices[link]

Depilation is the removal of hair from the surface of the skin. This can be achieved through methods such as shaving. Epilation is the removal of the entire hair strand, including the part of the hair that has not yet left the follicle. A popular way to epilate hair is through waxing it.

Shaving[link]

Many razors have multiple blades purportedly to ensure a close shave. While shaving initially will leave skin feeling smooth and hair free, new hair growth will appear a few hours after hair removal.

Shaving is accomplished with bladed instruments, such as razors. The blade is brought close to the skin and stroked over the hair in the desired area to cut the terminal hairs and leave the skin feeling smooth. Depending upon the rate of growth, one can begin to feel the hair growing back within hours of shaving. This is especially evident in men who develop a five o'clock shadow after having shaved their faces. This new growth is called stubble. Stubble typically appears to grow back thicker because the shaved hairs are blunted instead of tapered off at the end, although the hair never actually grows back thicker.

Waxing[link]

Waxing involves using a sticky wax and strip of paper or cloth to pull hair from the root. Waxing is the ideal hair removal technique to keep an area hair-free for long periods of time. It can take five to nine weeks for waxed hair to begin to resurface again. Hair in areas that have previously been waxed also is known to grow back finer and thinner, especially compared to hair that has been shaved with a razor.

Cutting and trimming[link]

Because the hair on the head is normally longer than other types of body hair, it is cut with scissors or clippers. People with longer hair will most often use scissors to cut their hair, whereas shorter hair is maintained using a trimmer. Depending on the desired length and overall health of the hair, periods without cutting or trimming the hair can vary.

Many people will confuse what a haircut is versus what a trim is. A haircut is usually performed in order to change one's hairstyle, while a trim helps to keep away split ends and keep the hair well-groomed. Cutting hair tends to take off more hair than trimming hair does. When hair is trimmed, only the first few centimeters need to be removed, whereas haircuts can sometimes result in the loss of many inches of hair.

Social role[link]

Portrait of a Woman, Alessandro Allori (1535–1607; Uffizi Gallery): a plucked hairline gives a fashionably "noble brow"

Hair has great social significance for human beings. It can grow on most external areas of the human body, except on the palms of the hands and the soles of the feet (among other areas). Hair is most noticeable on most people in a small number of areas, which are also the ones that are most commonly trimmed, plucked, or shaved. These include the face, ears, head, eyebrows, legs, and armpits, as well as the pubic region. The highly visible differences between male and female body and facial hair are a notable secondary sex characteristic.

Indication of status[link]

Healthy hair indicates health and youth (important in evolutionary biology). Hair colour and texture can be a sign of ethnic ancestry. Facial hair is a sign of puberty in men. White hair is a sign of age or genetics, which may be concealed with hair dye (not easily for some), although many prefer to assume it (specially if it is a poliosis characteristic of the person since childhood). Male pattern baldness is a sign of age, which may be concealed with a toupee, hats, or religious and cultural adornments. Although drugs and medical procedures exist for the treatment of baldness, many balding men simply shave their heads. Hair whorls have been discovered to be associated with brain development.

Hairstyle may be an indicator of group membership. During the English Civil War, the followers of Oliver Cromwell decided to crop their hair close to their head, as an act of defiance to the curls and ringlets of the king's men.^[43] This led to the Parliamentary faction being nicknamed Roundheads. Having bobbed hair was popular among the flappers in the 1920s as a sign of rebellion against traditional roles for women. Female art students known as the "cropheads" also adopted the style, notably at the Slade School in London, England. Regional variations in hirsutism cause practices regarding hair on the arms and legs to differ. Some religious groups may follow certain rules regarding hair as part of religious observance. The rules often differ for men and women.

Many subcultures have hairstyles which may indicate an unofficial membership. Many hippies, metalheads and Indian sadhus have long hair, as well many older indie kids. Many punks wear a hairstyle known as a mohawk or other spiked and dyed hairstyles; skinheads have short-cropped or completely shaved heads. Long stylized bangs were very common for emos, scene kids and younger indie kids in the 2000s and early 2010s, among people of both genders.

Heads were shaved in concentration camps, and head-shaving has been used as punishment, especially for women with long hair. The shaven head is common in military haircuts, while Western monks are known for the tonsure. By contrast, among some Indian holy men, the hair is worn extremely long.^{[citation needed]}

In the time of Confucius (5th century BCE), the Chinese grew out their hair and often tied it, as a symbol of filial piety.

Regular hairdressing in some cultures is considered a sign of wealth or status. The dreadlocks of the Rastafari movement were despised early in the movement's history. In some cultures, having one's hair cut can symbolize a liberation from one's past, usually after a trying time in one's life. Cutting the hair also may be a sign of mourning.

Tightly coiled hair in its natural state may be worn in an Afro. This hairstyle was once worn among African Americans as a symbol of racial pride. Given that the coiled texture is the natural state of some African Americans' hair, or perceived as being more "African", this simple style is now often seen as a sign of self-acceptance and an affirmation that the beauty norms of the (eurocentric) dominant culture are not absolute. It is important to note that African Americans as a whole have a variety of hair textures, as they are not an ethnically homogeneous group, but an ad-hoc of different racial admixtures.

The film Easy Rider (1969) includes the assumption that the two main characters could have their long hairs forcibly shaved with a rusty razor when jailed, symbolizing the intolerance of some conservative groups toward members of the counterculture.

At the conclusion of the Oz obscenity trials in the UK, the defendants had their heads shaved by the police, causing public outcry. During the appeal trial, they appeared in the dock wearing wigs.^{[citation needed]}

In the mid-2000s, a case where a 14-year-old student was prohibited of entering a school in Fortaleza ever again, according to him and his family because of his fauxhawk haircut, sparked nationwide polemics in Brazil.^[44][45] His family subsequently entered a court, and winned it in the early 2010s, receiving a reparation for constrant of R$3000 (which was originally R$10000), which is equivalent to roughly €1000-1100 or 1250-1350 U.S. dollars. Now he is 23, affirms to had acquired health problems related to high emotional stress and lives in San Francisco, California.^[45]

Religious practices[link]

Women's hair may be hidden using headscarves, a common part of the hijab in Islam and a symbol of modesty required for certain religious rituals in Orthodox Christianity. Russian Orthodox Church requires all married women to wear headscarves inside the church; this tradition is often extended to all women, regardless of marital status. Orthodox Judaism also commands the use of scarves and other head coverings for women for modesty reasons. Certain Hindu sects also wear head scarves for religious reasons. Sikhs generally keep their hair uncut and men keep it tied in a bun on the head, which is then covered appropriately using a turban. Multiple religions, both ancient and contemporary, require or advise one to allow their hair to become dreadlocks, though people also wear them for fashion. For men, Islam, Orthodox Judaism, Orthodox Christianity, Roman Catholicism and other religious groups have at various times recommended or required the covering of the head and sections of the hair of men, and some have dictates relating to the cutting of men's facial and head hair. Some Christian sects throughout history and up to modern times have also religiously proscribed the cutting of women's hair.

See also[link]

• Hypotrichosis, the state of having a less than normal amount of hair on the head or body
• Hypertrichosis, the state of having an excess of hair on the head or body
• Natural afro-hair
• Hair follicle
• Cowlick
• List of cutaneous conditions

References[link]

Notes[link]

This article needs additional citations for verification. Please help improve this article by adding citations to reliable sources. Unsourced material may be challenged and removed. (December 2007)

Bibliography[link]

• Iyengar, B. (1998). The hair follicle is a specialized UV receptor in human skin? Bio Signals Recep, 7(3), 188–194.
• Jablonski, N.G. (2006). Skin: a natural history. Berkley, CA: University of Califiornia Press.
• Rogers, Alan R.; Iltis, David & Wooding, Stephen (2004), “Genetic variation at the MC1R locus and the time since loss of human body hair”, Current Anthropology 45 (1): 105–108.
• Tishkoff, S.A. (1996). Global patterns of linkage disequilibrium at the CD4 locus and modern human origins. Science. 271(5254), 1380–1387.

External links[link]

Wikimedia Commons has media related to: Hair

Wikiquote has a collection of quotations related to: Hair

Look up hair in Wiktionary, the free dictionary.

• Discussion about shaving and cultures
• Answers to several questions related to hair from curious kids
• How to measure the diameter of your own hair using a laser pointer
• Instant insight outlining the chemistry of hair from the Royal Society of Chemistry

v t e Human regional anatomy (TA A01.1) Head Forehead · Ear · Jaw · Face (Cheek, Eye, Nose, Mouth, Chin) · Occiput · Scalp · Temple Neck Throat · Adam's apple Trunk Chest (Breast) · Abdomen (Navel) · Back · Pelvis (Sex organs) Limbs Upper limb Shoulder · Axilla · Arm Elbow · Forearm Wrist · Hand · Finger (Thumb · Index · Middle · Ring · Little) Lower limb/ (see also leg) Hip · Buttocks · Thigh Knee · Calf · Crus Ankle · Heel · Foot · Toe (Hallux · Long  • Third  • Fourth  • Fifth) · Sole General anatomy: systems and organs, regional anatomy, planes and lines, superficial axial anatomy, superficial anatomy of limbs

Kandi Burruss (born May 17, 1976), better known by her stage names Kandi or Kandi Girl, is an American R&B singer-songwriter, actress, record producer, and television personality. She is also a former member of the group Xscape from Atlanta.

Early life[link]

Kandi was born on May 17, 1976 in Atlanta, Georgia to a single mother, Joyce Jones^{[citation needed]}. Kandi has stated she was definitely a daddy's girl before her father left the family when Kandi was 4. Kandi was very close to her brother Patrick, who was older by 7 1/2 years and later died in a car accident when Kandi was a young teenager. Her first major appearance, on BET's Teen Summit, introduced her to the public. Kandi Attended Tri- Cities High School In East Point GA.

Career[link]

Songwriting[link]

After the disbanding of Xscape, Burruss began to focus on production and songwriting. In 1999, Burruss teamed with bandmate Tameka "Tiny" Cottle to score the international number one hit "No Scrubs" for TLC. The single received a Grammy for Best R&B Performance by a Duo or Group with Vocal, MTV Video Music Award, Billboard Music Award and Soul Train Music Award. That same year, Burruss scored a hit for Destiny's Child on their second album, "The Writing's on the Wall". "Bills, Bills, Bills", also topped the Billboard "Hot 100" chart & was followed by their second single "Bug a Boo", also penned by Burruss. Kandi would go on to work with then newcomer Pink by penning her debut single "There You Go" off Pink's debut album "Can't Take Me Home".

Burruss was the first African-American woman ever to win ASCAP's Songwriter of the Year award in 2000. She won for Songwriter of the year in the Rhythm & Soul category.^[1][2][3][4] She won for songwriting credits such as Destiny Child's "Bills, Bills, Bills" and TLC's No Scrubs.

Kandi's writing credits continued to grow by penning for the likes of Mariah Carey, Alicia Keys, Boyz II Men, Da Brat, NSYNC, N-Toon, Solo, Usher, Fantasia, Mýa, Joe, MC Lyte and Whitney Houston.

Kandi has since rewritten and executive produced "Tardy For The Party" for Real Housewives of Atlanta co-star Kim Zolciak's debut album. The song was made available to iTunes on September 1, 2009.

[edit] Kandi Koated and Reality Television (2009–present)

In early 2006, Kandi began work on her second album. The first single, "I Need", was a promotional single only. It featured rap duo 8Ball and MJG. Another song, "Keep It Gangsta" featuring rapper Lil Scrappy, was also set to appear on the album, although it was never released as a promo single.

Failing to generate any buzz, plans for the new album were temporarily put on hold. Burruss then teamed with Atlanta female rapper Rasheeda to form the duo Peach Candy, signed to D-Lo Entertainment. Their first single, titled "Bam", got little airplay and plans for this album were also put on hold. PeachCandy EP was released on iTunes with a 5-track lineup.

In 2009, Kandi joined the second season of the Bravo reality series, The Real Housewives of Atlanta, and went back to work on her second album, then titled B.L.O.G.. Her debut EP, the Fly Above EP, was released on October 29, 2009. The album will be released on her own record label, Kandi Koated Entertainment, and possibly Capitol Records, although the EP will only be released on Kandi Koated Entertainment. The album is confirmed to feature appearances from artists such as Rick Ross, Rasheeda, Gucci Mane and possibly Missy Elliott. Despite speculation that the lead single would be either "I Like Him" featuring Rick Ross and Rasheeda or "Trade Him In" featuring Gucci Mane, the lead single, "Fly Above", was released on October 6, 2009. It was later dubbed a promo single. Kandi signed a record deal with Asylum Records in early 2010^[5] and released her second album, Kandi Koated, on December 14, 2010. The lead single "Leave U" debuted at #89 on Billboard's Hot R&B/Hip-Hop Songs chart. Kandi also made a cameo appearance along with Atlanta producer Jermaine Dupri on the first episode in the new television series, Single Ladies.

Kandi is now working on her music and Bravo has greenlit her new reality show, The Kandi Factory. "The Kandi Factory will capture a unique look at her successful music career as Kandi transforms two music hopefuls into the next big pop star."^{[citation needed]}

Personal life[link]

Kandi briefly dated Gerald Levert.

Burruss and ex-boyfriend Russell "Block" Spencer of Block Entertainment had a daughter named Riley Spencer^[6] on August 22, 2002.^[7]

In late 2008, Kandi began a relationship with Ashley "A.J." Jewell and after dating several months, the couple became engaged to be married in January 2009. Tensions began to rise between Jewell, who participated in the second season of The Real Housewives of Atlanta, and Kandi's mother over Jewell's having six children of his own with four different mothers. In June 2009, Burruss and Jewell called off the engagement and later dissolved their romantic relationship completely, though Jewell was quoted as saying they remained "good friends". He moved out of Burruss' home, although his 12-year-old twin daughters remain in her care.

Less than four months after separating from Burruss, on October 2, 2009, Jewell was killed outside of a club's parking lot in a fight. He suffered massive head trauma in a heated argument at the Body Tap Club, an Atlanta strip club. The police arrested a club manager named Fredrick Richardson. He has been charged with voluntary manslaughter.^[8] However, sickle-cell anemia may have played a significant role in the death. According to the medical examiner, it is likely that Jewell was worked up from the fight and because of his condition, could not replenish the oxygen he lost during the incident, which the doctor says could have fatal consequences. Two days after Jewell's death, Kandi posted an open-letter expressing her heartfelt grief over her loss. In late 2010, Kandi released Kandi Koated. Her album includes tracks such as "Me and U", "Fly Above", and "Haven't Loved Right". Many songs on her album are inspired by her late ex-fiance AJ.

Kandi and three of her friends started a "sex and relationship" webshow on Ustream titled Kandi Koated Nights. They have had numerous guests, including Katt Stacks, Tionna T. Smalls, Grand Hustle, Jazze Pha, Young Joc. The show began to air on television beginning in 2011.^[9]

Discography[link]

• 2000: Hey Kandi
• 2010: Kandi Koated

Awards[link]

Grammy

• 2000 (42nd Grammy Awards)

• • Record of the Year – No Scrubs – TLC (Nominated)
  • Album of the Year FanMail – TLC (songwriter) (Nominated)
  • Best R&B Song (Rhythm & Blues Song) – TlC for "No Scrubs" (Won)
  • Best R&B Performance by a Duo or Group with Vocals – TLC for "No Scrubs"(Won)
  • Best R&B Song (Rhythm & Blues Song) – Destiny's Child for "Bills, Bills, Bills" (Nominated)
  • Best R&B Performance by a Duo or Group with Vocals – Destiny's Child for "Bills, Bills, Bills" (Nominated)

ASCAP

• 2000
  • Songwriter of the Year (Rhythm & Sou) for "Bills, Bills, Bills" and "No Scrubs" (Won)

References[link]

External links[link]

• Kandi Burruss Twitter
• Kandi official Myspace Page
• StraightFromTheA.com
• Kandi Burruss Singersroom.com
• Kandi Talks About Xscpae BreakUp, Ghettofabu.com
• Kaandi/Rasheeda Form Peach Candy Duo, Ballerstatus.com
• Peach Candy official Myspace Page
• Kandi Burruss Interview
• Interview, HitQuarters Mar 2007

v t e Xscape Kandi Burruss LaTocha Scott Tameka Scott Tameka "Tiny" Cottle Kiesha Miles Studio albums Hummin' Comin' at 'Cha Off the Hook Traces of My Lipstick Other albums Understanding Super Hits Singles "Just Kickin' It" "Understanding" "Love on My Mind" "Tonight" "Is My Living in Vain" "Feels So Good" "Who Can I Run To" "Do You Want To" "Can't Hang" "The Arms of the One Who Loves You" "My Little Secret" "What's Up" Related articles Discography Book:Xscape