The gizzard, also referred to as the ventriculus, gastric mill, and gigerium, is an organ found in the digestive tract of some animals, including birds, reptiles, earthworms and some fish. This specialized stomach constructed of thick, muscular walls is used for grinding up food; often rocks are instrumental in this process. In certain insects and mollusks, the gizzard features chitinous plates or teeth.
Etymology
The word "gizzard" comes from the
Middle English giser, which derives from a similar word in
Old French, which itself evolved from the
Latin gigeria, meaning giblets. The Latin word probably derives from a common
Proto-Indo-European source with the
Persian word for liver, which is "jigar".
Structure
, seen at the right of the
duodenum between the legs.]]
Birds swallow food and store it in their
crop if necessary. Then the food passes into their glandular stomach, also called the
proventriculus, which is also sometimes referred to as the true stomach. This is the secretory part of the stomach. Then the food passes into the ventriculus (also known as the
muscular stomach or
gizzard). The gizzard can grind the food with previously-swallowed stones and pass it back to the true stomach, and vice versa. Bird gizzards are lined with a tough layer made of the carbohydrate-protein complex
koilin, to protect the muscles in the gizzard and to aid in digestion.
Gizzard stones
Some animals that lack teeth will swallow stones or grit to aid in digestion. All birds have gizzards, but not all will swallow stones or grit. The birds that do, employ the following method of '
mastication':
:"
A bird swallows small bits of gravel that act as 'teeth' in the gizzard, breaking down hard food such as seeds and thus helping digestion." (Solomon et al., 2002).
These stones are called
gizzard stones or
gastroliths and are usually round and smooth from the polishing action in the animal's stomach. When too smooth to do their required work, they may be passed or regurgitated.
Animals with gizzards
Birds
All birds have gizzards. The gizzards of
emus,
turkeys,
chickens and
ducks are most notable in cuisine (see below).
Fish
The mullet (
Mugilidae) found in
estuarine waters worldwide, and the gizzard or mud
shad, found in freshwater lakes and streams from New York to Mexico, have gizzards.
The
gillaroo (
Salmo stomachius), a richly colored species of
trout found in the Irish lake,
lough Melvin in
County Fermangh, has a gizzard which is used to aid the digestion of water snails, the main component of its diet.
Reptiles
Alligators and
crocodiles also have gizzards.
Invertebrates
Most invertebrates also have gizzards. The gizzard is used to grind up food, and it is part of the digestive system.
Dinosaurs
Dinosaurs which are believed to have had gizzards based on the discovery of gizzard stones recovered near fossils include:
Psittacosaurus
Massospondylus
Sellosaurus
Omeisaurus
Apatosaurus
Barosaurus
Dicraeosaurus
Seismosaurus
Claosaurus had been believed to have a gizzard, but this was A)
Edmontosaurus annectens and B) probably based on stream-washed gravel; see Creisler, Benjamin S. 2007. Deciphering duckbills. Page 199 in Carpenter, Kenneth (ed.). Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs. Indiana University Press: Bloomington and Indianapolis.
In cuisine
Poultry gizzards are a popular food throughout the world. Grilled
chicken gizzards are sold as
street food in
Haiti and throughout
Southeast Asia. In
Indonesia, gizzard and liver are considered as part of a complete fried poultry dish. Stewed gizzards are eaten as a snack in
Portugal, while
pickled
turkey gizzards are a traditional food in some parts of the
Midwestern United States. In
Hungary it is made with
paprika. In
Nigeria gizzard is either grilled or fried and served with stew and fried plantain. In the Southern United States, the gizzard is typically served fried, sometimes eaten with hot sauce or honey mustard, or added to crawfish boil along with crawfish
sauce, and it is also used in traditional New Orleans
gumbo. In Chicago, gizzard is battered, deep fried and served with fries and sauce. Gizzard and mashed potato is a popular food in many European countries. In
France, especially the
Dordogne region, gizzards are eaten in the traditional Perigordian Salad, along with
walnuts,
croutons and
lettuce. The
Chamber of Commerce in
Potterville, Michigan have held a Gizzard Fest each June since 2000; a gizzard-eating contest is among the weekend's events.. In Iran some kebab restaurants mix chicken gizzards in their koobideh kebabs to increase the meat content.
The word "Sangdana" is commonly used to refer to chicken gizzards in Pakistan. The word is derived from Persian (Sang = stone and dana = grain). It may be served cooked in a curry while barbecued skewered gizzards are also popular.
In Yiddish, gizzards are referred to as "pipik'lach", literally meaning navels. The gizzards of kosher species of birds have a green or yellowish membrane lining the inside, which must be peeled off before cooking, as it lends a very bitter taste to the food. In traditional Eastern European Jewish cuisine, the gizzards, necks, and feet of chickens were often cooked together, although not the liver, which per Kosher law must be broiled. Kosher butchers often sell roasting chickens with the gizzard, neck, and feet butchered and left in the cavity to be used for making chicken soup.
In Gorkha cuisine, the gizzard and the liver are fried in oil with tomatoes, onions and chilies to prepare a popular sidedish. Called karchi-marchi, the dish is often an accompaniment to alcoholic drinks.
In Punjabi cuisine, the gizzard is broiled alongside mooli, chilli and garlic to produce a dish named 'Jib-Jab'. It is often served with prathar, which is dipped into the gizzard juice (the Jib-Jab juice).
In Uganda, Cameroon and Nigeria, the gizzard of a cooked chicken is traditionally set aside for the oldest or most respected male at the table.
Giblets consist of the heart, liver and gizzard of a bird, and are often eaten themselves or used as the basis for a soup or stock.
In Taiwan, gizzards are often slow-cooked and served hot or cold in slices, with green onions and soy sauce.
In Mainland China, duck gizzard is a common snack, eaten alongside other duck parts such as feet, neck, heart, tongue, or head. Areas famous for their gizzard are Sichuan and Hubei provinces. Wuhan city in Hubei is famous for its spicy gizzard, called Jiujiuya (Simplified Chinese:久久鸭). In Northern China, you can find BBQ duck gizzard.
Generic meaning
The term "gizzard" can also refer to the general guts, innards or entrails of animals.
References
Solomon, E.P., Berg L.P., and Martin D.W., 2002. Biology Sixth Edition. Thomson Learning Inc., Australia, Canada, Mexico, Singapore, Spain, United Kingdom, United States pp. 664
Dyce, Sack, Wensing, 2002. Textbook of Veterinary Anatomy Third Edition, Saunders. ISBN 0-7216-8966-3
Category:Animal anatomy
Category:Dinosaur anatomy
Category:Offal
Category:Bird anatomy