The common fruit fly, Drosophila melanogaster, has been the subject of much research

Invertebrates are animal species that do not develop a vertebral column. This in effect includes all animals apart from the subphylum Vertebrata. Familiar examples of invertebrates include insects, snails, and starfish. Taxonomically speaking "invertebrate" is no more than a term of convenience; only about 3% of animal species include a vertebral column in their anatomy.^[1] In other words all animals except those in the chordate subphylum Vertebrata (fish, amphibians, reptiles, birds, and mammals) are regarded as invertebrates. What is more, many an individual invertebrate taxon has a greater number and variety of species than the entire subphylum Vertebrata,^[2] In fact some of the so-called invertebrata, such as the Chaetognatha and Hemichordata, are more closely related the Chordata than to other invertebrate phyla.

Invertebrates form a massively paraphyletic group. It is generally accepted that the phyla comprising modern Eukaryota share a common multicellular ancestor, but with the sole exception of one subphylum of the phylum Chordata, all those phyla are classed as invertebrates along with two of the three subphyla in the Chordata: Tunicata and Cephalochordata. These two, plus all the other known invertebrates, have only one cluster of Hox genes, while the vertebrates have duplicated their original cluster more than once.^{[citation needed]}

Within palaeozoology and palaeobiology, invertebrates are often studied within the fossil discipline called invertebrate palaeontology.

Etymology[link]

The word invertebrate comes from the word vertebrate, with the prefix in- attached to it.^[3] Thus, the word indicates that which lacks vertebrae.

Taxonomic significance[link]

The term Invertebrate promotes a great deal of confusion among non-biologists. It does not refer to any particular taxon in the same way that Arthropoda, Vertebrata or Manidae would do. Each of those examples describes a presumably monophyletically valid taxon, say a phylum, subphylum or family. In referring to taxonomy of the Animalia, "invertebrata" is a term of convenience, not a taxon; it has very little circumscriptional significance except arguably within the Chordata. The Vertebrata as a subphylum comprises such a small proportion of the Metazoa that to speak of the kingdom Animalia in terms of "Vertebrata" and "Invertebrata" would be about as practical as classifying animals into mayflies and non-mayflies, or transport into rowing boats and non-rowing boats. It would be logically correct to do so, and rowing boats as such do form a practical group, but speaking of "non-rowing boats" would lump together land, sea, air and space transport in ways that rarely would be useful. In formal taxonomy of the Animalia there are higher level attributes that logically should precede the presence or absence of the vertebral column in constructing a cladogram, for example, the presence of a notochord. That would at least circumscribe the Chordata. However, even the notochord would be a less fundamental criterion than aspects of embryological development and symmetry^[4] or perhaps bauplan.^[5] The resultant cladistic structure would not resemble anything like a binary split into vertebrates and invertebrates.

At the same time, there certainly is no reason to avoid the use of the terms Invertebrata or invertebrates when they are convenient, but it is important not to confuse the terms with the names or relationships of actual taxa.

Characteristics[link]

The trait that is common to all invertebrates is the absence of a vertebral column: this creates a distinction between invertebrates and vertebrates. Being animals, invertebrates are heterotrophs, and require sustenance in the form of the consumption of other organisms. With a few exceptions, such as the Porifera, invertebrates generally have bodies composed of differentiated tissues. There is also typically a digestive chamber with one or two openings to the exterior.

Like vertebrates, most invertebrates reproduce at least partly through sexual reproduction. They produce specialized reproductive cells that undergo meiosis to produce smaller, motile spermatozoa or larger, non-motile ova.^[6] These fuse to form zygotes, which develop into new individuals.^[7] Others are capable of asexual reproduction, or sometimes, both methods of reproduction.

Phyla[link]

The fossil coral Cladocora from the Pliocene of Cyprus

The term invertebrate covers several phyla. One of these are the sponges (Porifera). They were long thought to have diverged from other animals early.^[8] They lack the complex organization found in most other phyla.^[9] Their cells are differentiated, but in most cases not organized into distinct tissues.^[10] Sponges typically feed by drawing in water through pores.^[11] Some speculate that sponges are not so primitive, but may instead be secondarily simplified.^[12] The Ctenophora and the Cnidaria, which includes sea anemones, corals, and jellyfish, are radially symmetric and have digestive chambers with a single opening, which serves as both the mouth and the anus.^[13] Both have distinct tissues, but they are not organized into organs.^[14] There are only two main germ layers, the ectoderm and endoderm, with only scattered cells between them. As such, they are sometimes called diploblastic.^[15]

The Echinodermata are radially symmetric and exclusively marine, including starfish (Asteroidea), sea urchins, (Echinoidea), brittle stars (Ophiuroidea), sea cucumbers (Holothuroidea) and feather stars (Crinoidea).^[16]

Other phyla of invertebrates are the Hemichordata, or acorn worms,^[17] and the Chaetognatha, or arrow worms.

The largest animal phylum is also included within invertebrates: the Arthropoda, including insects, spiders, crabs, and their kin. All these organisms have a body divided into repeating segments, typically with paired appendages. In addition, they possess a hardened exoskeleton that is periodically shed during growth.^[18] Two smaller phyla, the Onychophora and Tardigrada, are close relatives of the arthropods and share these traits. The Nematoda or roundworms, are perhaps the second largest animal phylum, and are also invertebrates. Roundworms are typically microscopic, and occur in nearly every environment where there is water.^[19] A number are important parasites.^[20] Smaller phyla related to them are the Kinorhyncha, Priapulida, and Loricifera. These groups have a reduced coelom, called a pseudocoelom. Other invertebrates include the Nemertea or ribbon worms, and the Sipuncula.

Another phylum is Platyhelminthes, the flatworms.^[21] These were originally considered primitive, but it now appears they developed from more complex ancestors.^[22] Flatworms are acoelomates, lacking a body cavity, as are their closest relatives, the microscopic Gastrotricha.^[23] The Rotifera or rotifers, are common in aqueous environments. Invertbrates also include the Acanthocephala or spiny-headed worms, the Gnathostomulida, Micrognathozoa, and the Cycliophora.^[24]

Also included are two of the most successful animal phyla, the Mollusca and Annelida.^[25][26] The former, which is the second-largest animal phylum by number of described species, includes animals such as snails, clams, and squids, and the latter comprises the segmented worms, such as earthworms and leeches. These two groups have long been considered close relatives because of the common presence of trochophore larvae, but the annelids were considered closer to the arthropods because they are both segmented.^[27] Now, this is generally considered convergent evolution, owing to many morphological and genetic differences between the two phyla.^[28]

Other phyla include Acoelomorpha, Brachiopoda, Bryozoa, Entoprocta, Phoronida, and Xenoturbellida.

History[link]

Some of the first animal fossils appear to be those of invertebrates. 665-million-year-old fossils in the Trezona Formation at Trezona Bore, West Central Flinders, South Australia have been interpreted as being early sponges.^[29] Some paleontologists suggest that animals appeared much earlier, possibly as early as 1 billion years ago.^[30] Trace fossils such as tracks and burrows found in the Tonian era indicate the presence of triploblastic worms, like metazoans, roughly as large (about 5 mm wide) and complex as earthworms.^[31]

Around 453 MYA, animals began diversifying, and many of the important groups of invertebrates diverged from one another. Fossils of invertebrates are found in various types of sediment from the Phanerozoic.^[32] Fossils of invertebrates are commonly used in stratigraphy.^[33]

Classification[link]

Carl Linnaeus divided these animals into only two groups, the Insecta and the now-obsolete Vermes (worms). Jean-Baptiste Lamarck, who was appointed to the position of "Curator of Insecta and Vermes" at the Muséum National d'Histoire Naturelle in 1793, both coined the term "invertebrate" to describe such animals, and divided the original two groups into ten, by splitting Arachnida and Crustacea from the Linnean Insecta, and Mollusca, Annelida, Cirripedia, Radiata, Coelenterata and Infusoria from the Linnean Vermes. They are now classified into over 30 phyla, from simple organisms such as sea sponges and flatworms to complex animals such as arthropods and molluscs.

Significance of the group[link]

Invertebrates are animals without a vertebral column. This has led to the conclusion that invertebrates are a group that deviates from the norm, vertebrates. This has been said to be due to the fact that researchers in the past, such as Lamarck, viewed vertebrates as a "standard": in Lamarck's theory of evolution, he believed that characteristics acquired through the evolutionary process involved not only survival, but also progression toward a "higher form", to which humans and vertebrates were closer than invertebrates were. Although goal-directed evolution has been abandoned, the distinction of invertebrates and vertebrates persists to this day, even though the grouping has been noted to be "hardly natural or even very sharp." Another reason cited for this continued distinction is that Lamarck created a precedent through his classifications which is now difficult to escape from. It's also possible that some humans believe that, they themselves being vertebrates, the group deserves more attention than invertebrates.^[34] In any event, in the 1968 edition of Invertebrate Zoology, it is noted that "division of the Animal Kingdom into vertebrates and invertebrates is artificial and reflects human bias in favor of man's own relatives." The book also points out that the group lumps a vast number of species together, so that no one characteristic describes all invertebrates. In addition, some species included are only remotely related to one another, with some more related to vertebrates than other invertebrates.^[35]

In research[link]

Two of the most commonly studied model organisms are invertebrates: the fruit fly Drosophila melanogaster and the nematode Caenorhabditis elegans. They have long been the most intensively studied model organisms, and were among the first life-forms to be genetically sequenced. This was facilitated by the severely reduced state of their genomes, but many genes, introns, and linkages have been lost. Analysis of the starlet sea anemone genome has emphasised the importance of sponges, placozoans, and choanoflagellates, also being sequenced, in explaining the arrival of 1500 ancestral genes unique to animals.^[36]

Arthropods, especially insects, are often used by forensic scientists. For example, some invertebrates are attracted to dead bodies.^[18]

See also[link]

• Marine invertebrates
• Pain in invertebrates

References[link]

Further reading[link]

• Hyman, L. H. 1940. The Invertebrates (6 volumes) New York : McGraw-Hill. A classic work.
• Anderson, D. T. (Ed.). (2001). Invertebrate zoology (2nd ed.). Oxford: Oxford University Press.
• Brusca, R. C., & Brusca, G. J. (2003). Invertebrates (2nd ed.). Sunderland, Mass. : Sinauer Associates.
• Miller, S.A., & Harley, J.P. (1996). Zoology (4th ed.). Boston: WCB/McGraw-Hill.
• Pechenik, Jan A. (2005). Biology of the invertebrates. Boston: McGraw-Hill, Higher Education. pp. 590 pp. ISBN 0-07-234899-2. 
• Ruppert, E. E., Fox, R. S., & Barnes, R. D. (2004). Invertebrate zoology: a functional evolutionary approach. Belmont, CA: Thomas-Brooks/Cole.

External links[link]

• A. R. Maggenti & S. Gardner (2005). Online Dictionary of Invertebrate Zoology. http://digitalcommons.unl.edu/onlinedictinvertzoology/. 
• Support for endangered invertebrates
• African Invertebrates

Bill Nye
Nye at Bridgewater State University in 2007
Born	(1955-11-27) November 27, 1955 (age 56)[1] Washington, D.C.[1]
Residence	Los Angeles
Fields	Mechanical engineering
Institutions	Boeing Cornell University Planetary Society
Alma mater	Cornell University (BS)
Known for	Bill Nye the Science Guy
Signature

William Sanford "Bill" Nye (born November 27, 1955),^[2] popularly known as Bill Nye the Science Guy, is an American science educator, comedian, television host, actor, mechanical engineer, and scientist. He is best known as the host of the Disney/PBS children's science show Bill Nye the Science Guy (1993–1998) and for his many subsequent appearances in popular media as a science educator.

Early life and career[link]

William Sanford Nye was born in Washington, D.C., the son of Jacqueline (née Jenkins; c. 1920–2000), a codebreaker during World War II, and Edwin Darby "Ned" Nye (died 1997), also a World War II veteran whose experience in a Japanese prisoner of war camp led him to become a sundial enthusiast.^[3][4][5] Nye is a fourth-generation Washington, D.C. resident on his father's side of the family. After attending Lafayette Elementary and Alice Deal Junior High in the city, he was accepted to the private Sidwell Friends School on a partial scholarship, graduating in 1973.^[6][7] He studied mechanical engineering at Cornell University, where one of his professors was Carl Sagan,^[8] and graduated with a Bachelor of Science in 1977.^[9] He was awarded an honorary doctorate by The Johns Hopkins University in May 2008. In May 2011, Nye was awarded an Honorary Doctor of Science degree from Willamette University where he was the keynote speaker for that year's commencement exercises.^[10]

Nye began his career in Seattle at Boeing, where, among other things, he starred in training films and developed a hydraulic pressure resonance suppressor still used in the 747. Later he worked as a consultant in the aeronautics industry. Nye told the St. Petersburg Times in 1999 that he applied to be a NASA astronaut every few years but was always rejected.^[11]

The Science Guy[link]

Nye began his professional entertainment career as an actor on a local sketch comedy television show in Seattle, Almost Live!; Nye attempted to correct the host of Almost Live! after the host pronounced "gigawatt" as "jigowatt."^[12][13] The character name came from the host's comment "Who do you think you are—Bill Nye the Science Guy?" and Nye was thereafter known as such on the program. His other main recurring role on Almost Live! was as Speedwalker, a speedwalking Seattle superhero.

Coincidentally, in 1991–93 he appeared in the live-action educational segments of Back to the Future: The Animated Series with a nonspeaking role as an assistant to Dr. Emmett Brown, played by Christopher Lloyd, in which he would demonstrate science with Lloyd's voice-over. The segments' national popularity led to Nye's hosting the educational television program Bill Nye the Science Guy from 1993 to 1998. Each of the 100 episodes aimed to teach a specific topic in science to a preteen audience, yet it garnered a wide adult audience as well. The show was very popular as a school resource and is still used to this day. He has written several books as The Science Guy. In addition to hosting the show, he was also a writer and producer for the series, which was filmed entirely in Seattle.

When portraying "the Science Guy", Nye wears a light blue lab coat and a bow tie and takes on the persona of an excited, jocular science educator. This popular image of Nye has been parodied by numerous sources, including the webcomic xkcd^[14] and the satirical news organization The Onion.^[15][16] In response to the fake headline "Crack Nearly Killed Me," Nye took the joke in good humor and sent The Onion an email thanking them for "dealing compassionately with this matter."^[17]

He appeared in his Science Guy persona alongside Ellen DeGeneres and Alex Trebek in Ellen's Energy Adventure, an attraction that has played since 1996 at the Universe of Energy pavilion inside Epcot at Walt Disney World. He also has a voice-over at the DINOSAUR attraction in Disney's Animal Kingdom park, where he tells guests about the dinosaurs while they queue for the ride. In addition, he appears in the "Design Lab" of CyberSpace Mountain inside DisneyQuest at Walt Disney World, where he refers to himself as "Bill Nye the Coaster Guy."

[edit] Post-Science Guy career

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Bill Nye "The Science Guy" and Executive Director of The Planetary Society received the Committee for Skeptical Inquiry's "In Praise of Reason" Award at CSICON 2011 in New Orleans

Nye remained interested in science education through entertainment. He created a 13-episode PBS KCTS-TV series about science, called The Eyes of Nye, aimed at an older audience than his previous show had been. Airing in 2005, it often featured episodes based on politically relevant themes such as genetically modified food, global warming, and race.

He played a science teacher in Disney's 1998 TV movie The Principal Takes a Holiday; he made a hovercraft in order to demonstrate science in an unusual classroom manner. From 2000 to 2002, Nye was the technical expert in BattleBots. In 2004 and 2005, Nye hosted 100 Greatest Discoveries, an award-winning series produced by THINKFilm for The Science Channel and in high definition on the Discovery HD Theater. He was also host of an eight-part Discovery Channel series called Greatest Inventions with Bill Nye.

Nye has guest-starred in several episodes of the crime drama Numb3rs as an engineering faculty member. A lecture Nye gave several years ago on exciting children about math was an inspiration for creating the Numb3rs show.^[18]

Bill joined the American Optometric Association in a multimedia advertising campaign to educate parents on the importance of getting their children a comprehensive eye examination.^[19]

He was a regular in TV Land discussions. He has also made guest appearances on the VH1 reality show America's Most Smartest Model.^[20]

Nye appears in segments of The Climate Code on The Weather Channel, telling his personal ways of saving energy. He still makes regular appearances on the show, often asking quiz questions.

As of fall 2008, Nye also appears on the daytime game show Who Wants to Be a Millionaire as part of the show's reintroduced "Ask the Expert" lifeline. In 2008, he also hosted Stuff Happens, a show on the then new Planet Green network.

In November 2008, Nye appeared in an acting role as himself in the fifth-season episode "Brain Storm" of Stargate Atlantis alongside fellow television personality and astrophysicist Neil deGrasse Tyson.^[21]

Nye has appeared numerous times on the talk show Larry King Live, speaking about topics such as global warming^[22] and UFOs.^[23] He argued that global warming is an issue that should be addressed by governments of the world in part because it could be implicated in the record-setting 2005 Atlantic hurricane season. On UFOs he has been skeptical of extraterrestrial explanations for sightings such as those at Roswell and Malmstrom Air Force Base in 1967.

In 2009, portions of Bill Nye's shows were used as lyrics and portions of the second Symphony of Science science education music video by composer John Boswell.

Nye recorded a short YouTube video (as himself, not his TV persona) advocating clean energy climate change legislation on behalf of Al Gore's Repower America campaign in October 2009.^[24]

Nye (as his TV persona) also made a guest appearance on The Dr. Oz Show.

On March 12, 2011, Nye made an appearance on CNN to discuss the evolving nuclear incidents in Japan as a result of the devastating earthquake and tsunami there. Nye erroneously stated that cesium is used to "slow and control" the nuclear reaction. ^[25] In reality, cesium (specifically cesium-137) is a nuclear fission product, not a control rod material. Nye also erroneously stated that the nuclear reactor involved in the Three Mile Island incident is still running and that the use of boron to slow the nuclear chain reaction is uncommon, when in fact boron-10 is commonly used in control rods, and is circulated in the coolant of reactors in the United States, as well as stored on site as a method of emergency shutdown.^[26][27]

A solar noon clock atop Rhodes Hall was gifted from Nye to Cornell on Aug. 27 following a public lecture that filled the 715-seat Statler Auditorium. Nye talked about his father's passion for sundials and timekeeping, his time at Cornell, his work on the sundials mounted on the Mars rovers and the story behind the Bill Nye Solar Noon Clock. ^[28]

Life outside television[link]

Nye speaking in 2010.

In the early 2000s, Nye assisted in the development of a small sundial that was included in the Mars Exploration Rover missions. Known as MarsDial, it included small colored panels to provide a basis for color calibration in addition to helping keep track of time.^[29] From 2005 to 2010 Nye was the vice president of The Planetary Society, an organization that advocates space science research and the exploration of other planets, particularly Mars.^[30] He became the organization's second Executive Director in September 2010 when Louis Friedman stepped down.^[31][32]

In November 2010, Nye became the face of a new permanent exhibition at the Chabot Space & Science Center in Oakland, CA. Bill Nye’s Climate Lab features Nye as commander of the Clean Energy Space Station, and invites visitors on an urgent mission to thwart climate change. Beginning with a view of Planet Earth from space, visitors explore air, water, and land galleries to discover how climate change affects Earth’s connected systems, and how to use the Sun, wind, land, and water to generate clean energy. In an interview about the exhibit, Nye said, “Everything in the exhibit is geared to showing you that the size of the problem of climate change is big. Showing you a lot about energy use... It’s a huge opportunity... We need young people, entrepreneurs, young inventors, young innovators to change the world.” ^[33]

He holds several United States patents,^[34] including one for ballet pointe shoes^[30] and another for an educational magnifying glass created by filling a clear plastic bag with water.^[35][36] From 2001–2006 Nye served as Frank H.T. Rhodes Class of '56 University Professor at Cornell University.^[9][37] Nye supported the 2006 reclassification of Pluto from a planet to a dwarf planet by the International Astronomical Union.^[38]

Since 2006, Nye has lived in Los Angeles in a house with modifications, though has owned a house on Mercer Island.^{[citation needed]} As of July 2007, Nye and environmental activist Ed Begley, Jr. are engaging in a friendly competition "to see who could have the lowest carbon footprint," according to Begley.^[39] In a 2008 interview, Nye joked that he wants to "crush Ed Begley" in their environmental competition.^[40] Nye and Begley are neighbors in Los Angeles, and sometimes dine together at a local vegetarian restaurant.^[40] Nye often appears on Begley's Planet Green reality show Living with Ed.

Nye made an appearance in Palmdale's 2010 video "Here Comes the Summer"^[41]; the band's lead singer Kay Hanley is his neighbor.

Nye is a fellow of the Committee for Skeptical Inquiry, a U.S. non-profit scientific and educational organization whose aim is to promote scientific inquiry, critical investigation, and the use of reason in examining controversial and extraordinary claims.^[42]

Interviewed by John Rael for the Independent Investigation Group IIG, Nye stated that his "concern right now... scientific illiteracy... you don't [the public] have enough rudimentary knowledge of the universe to evaluate claims."^[43]

Personal life[link]

Nye announced his engagement during an appearance on The Late Late Show with Craig Ferguson and was married briefly to his fiancée of five months, author Blair Tindall, on February 3, 2006. The ceremony was performed by Rick Warren at The Entertainment Gathering at the Skirball Cultural Center in Los Angeles. Yo-Yo Ma provided the music.^[44] Nye left the relationship seven weeks later when the marriage license was declared invalid;^[45] their impromptu ceremony, which preceded the license purchase, violated California state law, said Tindall in a radio interview.^[46] Nye later secured a temporary restraining order against Tindall after she entered his property late at night to pour weed killer in his garden.^[45]

Nye enjoys baseball and occasionally does experiments involving the physics of the game. As a longtime Seattle resident before becoming an entertainer, he is said to have been a fan of the Seattle Mariners, although recently he has voiced his preference (as a D.C. native) for the Washington Nationals.^[6]

References[link]

External links[link]

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Biography portal

Wikimedia Commons has media related to: Bill Nye

• Official website
• Bill Nye at the Internet Movie Database
• Bill Nye on Twitter
• Nye-Tindall wedding video
• An interview/discussion with Nye as guest on the radio show Loveline
• "Changing The World With Science Education" interview on Point of Inquiry.
• Seattle Times interview (April 26, 2004)
• Science Channel interview

Danny Baranowsky (also Danny B or dB Soundworks) is an electronic music composer, composing music mainly for indie films and indie games.^[2] He founded dB Soundworks to sell and promote his music. He is most known for providing the sound track for games Super Meat Boy,^[3][4] Canabalt^[1] and The Binding of Isaac.^[5]

References[link]

External links[link]

• dB Soundworks home page

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