The Paleolithic (or Palæolithic) Age, Era or Period, is a prehistoric period of human history distinguished by the development of the most primitive stone tools discovered (Modes I and II), and covers roughly 99% of human technological prehistory. It extends from the earliest known use of stone tools, probably by Hominins such as Australopithecines, 2.6 million years ago, to the end of the Pleistocene around 10,000 BP.^[1] The Paleolithic era is followed by the Mesolithic. The date of the Paleolithic—Mesolithic boundary may vary by locality as much as several thousand years.

During the Paleolithic, humans grouped together in small societies such as bands, and subsisted by gathering plants and hunting or scavenging wild animals.^[2] The Paleolithic is characterized by the use of knapped stone tools, although at the time humans also used wood and bone tools. Other organic commodities were adapted for use as tools, including leather and vegetable fibers; however, due to their nature, these have not been preserved to any great degree. Surviving artifacts of the Paleolithic era are known as paleoliths. Humankind gradually evolved from early members of the genus Homo such as Homo habilis — who used simple stone tools — into fully behaviorally and anatomically modern humans (Homo sapiens sapiens) during the Paleolithic era.^[3] During the end of the Paleolithic, specifically the Middle and or Upper Paleolithic, humans began to produce the earliest works of art and engage in religious and spiritual behavior such as burial and ritual.^[2][4][5][6] The climate during the Paleolithic consisted of a set of glacial and interglacial periods in which the climate periodically fluctuated between warm and cool temperatures.

The term Paleolithic was coined by archaeologist John Lubbock in 1865. It derives from Greek: παλαιός, palaios, "old"; and λίθος, lithos, "stone", literally meaning "old age of the stone" or "Old Stone Age."

One rich source of Paleothic artifacts has been the Euphrates river valley. Excavations started in the 1960s, when the Turkish government built the Keban dam on the river. The Keban historical salvage project was organized by Kemal Kurdas, then rector of Middle East Technical University, and a team of Turkish, American and Dutch archeologists led by Maurits van Loon excavated. Later more dams were built and salvage operations took place, unearthing settlements going back to the Paleolithic.

Human evolution[link]

This cranium, of Homo heidelbergensis, a Lower Paleolithic predecessor to Homo neanderthalensis and possibly Homo sapiens, dates to sometime between 500,000 to 400,000 BP.

Human evolution is the part of biological evolution concerning the emergence of humans as a distinct species.

Paleogeography and climate[link]

The Paleolithic climate consisted of a set of glacial and interglacial periods

The climate of the Paleolithic Period spanned two geologic epochs known as the Pliocene and the Pleistocene. Both of these epochs experienced important geographic and climatic changes that affected human societies.

During the Pliocene, continents continued to drift from possibly as far as 250 km from their present locations to positions only 70 km from their current location. South America became linked to North America through the Isthmus of Panama, bringing a nearly complete end to South America's distinctive marsupial fauna. The formation of the Isthmus had major consequences on global temperatures, because warm equatorial ocean currents were cut off, and the cold Arctic and Antarctic waters lowered temperatures in the now-isolated Atlantic Ocean. Central America formed completely during the Pliocene, allowing fauna from North and South America to leave their native habitats and colonize new areas.^[7] Africa's collision with Asia created the Mediterranean Sea, cutting off the remnants of the Tethys Ocean. During the Pleistocene, the modern continents were essentially at their present positions; the tectonic plates on which they sit have probably moved at most 100 km from each other since the beginning of the period.^[8]

Climates during the Pliocene became cooler and drier, and seasonal, similar to modern climates. Ice sheets grew on Antarctica. The formation of an Arctic ice cap around 3 Ma is signaled by an abrupt shift in oxygen isotope ratios and ice-rafted cobbles in the North Atlantic and North Pacific ocean beds.^[9] Mid-latitude glaciation probably began before the end of the epoch. The global cooling that occurred during the Pliocene may have spurred on the disappearance of forests and the spread of grasslands and savannas.^[7]

The Pleistocene climate was characterized by repeated glacial cycles during which continental glaciers pushed to the 40th parallel in some places. Four major glacial events have been identified, as well as many minor intervening events. A major event is a general glacial excursion, termed a "glacial". Glacials are separated by "interglacials". During a glacial, the glacier experiences minor advances and retreats. The minor excursion is a "stadial"; times between stadials are "interstadials". Each glacial advance tied up huge volumes of water in continental ice sheets 1500–3000 m deep, resulting in temporary sea level drops of 100 m or more over the entire surface of the Earth. During interglacial times, such as at present, drowned coastlines were common, mitigated by isostatic or other emergent motion of some regions.

Many great mammals such as wooly mammoths, wooly rhinoceros, and cave lions inhabited places like Siberia during the Pleistocene.

Paleoindians hunting a glyptodont. Glyptodonts were hunted to extinction within 2 millennia after humans' arrival to South America.

The effects of glaciation were global. Antarctica was ice-bound throughout the Pleistocene and the preceding Pliocene. The Andes were covered in the south by the Patagonian ice cap. There were glaciers in New Zealand and Tasmania. The now decaying glaciers of Mount Kenya, Mount Kilimanjaro, and the Ruwenzori Range in east and central Africa were larger. Glaciers existed in the mountains of Ethiopia and to the west in the Atlas mountains. In the northern hemisphere, many glaciers fused into one. The Cordilleran ice sheet covered the North American northwest; the Laurentide covered the east. The Fenno-Scandian ice sheet covered northern Europe, including Great Britain; the Alpine ice sheet covered the Alps. Scattered domes stretched across Siberia and the Arctic shelf. The northern seas were frozen. During the late Upper Paleolithic (Latest Pleistocene) c. 18,000 BP, the Beringa land bridge between Asia and North America was blocked by ice,^[8] which may have prevented early Paleo-Indians such as the Clovis culture from directly crossing Beringa to reach the Americas.

According to Mark Lynas (through collected data), the Pleistocene's overall climate could be characterized as a continuous El Niño with trade winds in the south Pacific weakening or heading east, warm air rising near Peru, warm water spreading from the west Pacific and the Indian Ocean to the east Pacific, and other El Niño markers.^[10]

The ice age ended with the end of the Paleolithic era (the end of the Pleistocene epoch), and Earth's climate became warmer. This may have caused or contributed to the extinction of the Pleistocene megafauna, although it is also possible that the late Pleistocene extinctions were (at least in part) caused by other factors such as disease and over hunting by humans.^[11][12] New research suggests that the extinction of the woolly mammoth may have been caused by the combined effect of climatic change and human hunting.^[12] Scientists suggest that climate change during the end of the Pleistocene caused the mammoths' habitat to shrink in size, resulting in a drop in population. The small populations were then hunted out by Paleolithic humans.^[12] The global warming that occurred during the end of the Pleistocene and the beginning of the Holocene may have made it easier for humans to reach mammoth habitats that were previously frozen and inaccessible.^[12] Small populations of wooly mammoths survived on isolated Arctic islands, Saint Paul Island and Wrangel Island, till circa 3700 and 1700 BCE respectively. The Wrangel Island population went extinct around the same time the island was settled by prehistoric humans.^[13] There's no evidence of prehistoric human presence on Saint Paul island (though early human settlements dating as far back as 6500 BCE were found on nearby Aleutian Islands).^[14]

Human way of life[link]

An artist's rendering of a temporary wood house, based on evidence found at Terra Amata (in Nice, France) and dated to the Lower Paleolithic (c. 400,000 BP)

Due to a lack of written records from this time period, nearly all of our knowledge of Paleolithic human culture and way of life comes from archaeology and ethnographic comparisons to modern hunter-gatherer cultures such as the Kung San who live similarly to their Paleolithic predecessors.^[16] The economy of a typical Paleolithic society was a hunter-gatherer economy.^[17] Humans hunted wild animals for meat and gathered food, firewood, and materials for their tools, clothes, or shelters.^[17] Human population density was very low, around only one person per square mile.^[2] This was most likely due to low body fat, infanticide, women regularly engaging in intense endurance exercise,^[18] late weaning of infants and a nomadic lifestyle.^[2] Like contemporary hunter-gatherers, Paleolithic humans enjoyed an abundance of leisure time unparalleled in both Neolithic farming societies and modern industrial societies.^[17][19] At the end of the Paleolithic, specifically the Middle and or Upper Paleolithic, humans began to produce works of art such as cave paintings, rock art and jewellery and began to engage in religious behavior such as burial and ritual.^[20]

Technology[link]

Picture of two Lower Paleolithic bifaces

Picture of a stone ball from a set of Paleolithic bolas

Paleolithic humans made tools of stone, bone, and wood.^[17] The earliest Paleolithic stone tool industry, the Olduwan, was developed by the earliest members of the genus Homo such as Homo habilis, around 2.6 million years ago.^[21] It contained tools such as choppers, burins and awls. It was completely replaced around 250,000 years ago by the more complex Acheulean industry, which was first conceived by Homo ergaster around 1.8 or 1.65 million years ago.^[22] The most recent Lower Paleolithic (Acheulean) implements completely vanished from the archeological record around 100,000 years ago and were replaced by more complex Middle Paleolithic/Middle Stone Age tool kits such as the Mousterian and the Aterian industries.^[23]

Lower Paleolithic humans used a variety of stone tools, including hand axes and choppers. Although they appear to have used hand axes often, there is disagreement about their use. Interpretations range from cutting and chopping tools, to digging implements, flake cores, the use in traps and a purely ritual significance, maybe in courting behavior. William H. Calvin has suggested that some hand axes could have served as "killer Frisbees" meant to be thrown at a herd of animals at a water hole so as to stun one of them. There are no indications of hafting, and some artifacts are far too large for that. Thus, a thrown hand axe would not usually have penetrated deeply enough to cause very serious injuries. Nevertheless, it could have been an effective weapon for defense against predators. Choppers and scrapers were likely used for skinning and butchering scavenged animals and sharp ended sticks were often obtained for digging up edible roots. Presumably, early humans used wooden spears as early as 5 million years ago to hunt small animals, much as their relatives, chimpanzees, have been observed to do in Senegal, Africa.^[24] Lower Paleolithic humans constructed shelters such as the possible wood hut at Terra Amata.

Fire was used by the Lower Paleolithic hominid Homo erectus/Homo ergaster as early as 300,000 or 1.5 million years ago and possibly even earlier by the early Lower Paleolithic (Oldowan) hominid Homo habilis and/or by robust australopithecines such as Paranthropus.^[2] However, the use of fire only became common in the societies of the following Middle Stone Age/Middle Paleolithic Period.^[1] Use of fire reduced mortality rates and provided protection against predators.^[25] Early hominids may have begun to cook their food as early as the Lower Paleolithic (c. 1.9 million years ago) or at the latest in the early Middle Paleolithic (c. 250,000 years ago).^[26] Some scientists have hypothesized that Hominids began cooking food to defrost frozen meat, which would help ensure their survival in cold regions.^[26]

The Lower Paleolithic hominid Homo erectus possibly invented rafts (c. 800,000 or 840,000 BP) to travel over large bodies of water, which may have allowed a group of Homo erectus to reach the island of Flores and evolve into the small hominid Homo floresiensis. However, this hypothesis is disputed within the anthropological community.^[27][28][29] The possible use of rafts during the Lower Paleolithic may indicate that Lower Paleolithic Hominids such as Homo erectus were more advanced than previously believed, and may have even spoken an early form of modern language.^[28] Supplementary evidence from Neanderthal and Modern human sites located around the Mediterranean Sea such as Coa de sa Multa (c. 300,000 BP) has also indicated that both Middle and Upper Paleolithic humans used rafts to travel over large bodies of water (i.e. the Mediterranean Sea) for the purpose of colonizing other bodies of land.^[30][28]

Around 200,000 BP, Middle Paleolithic Stone tool manufacturing spawned a tool making technique known as the prepared-core technique, that was more elaborate than previous Acheulean techniques.^[3] This technique increased efficiency by allowing the creation of more controlled and consistent flakes.^[3] It allowed Middle Paleolithic humans to create stone tipped spears, which were the earliest composite tools, by hafting sharp, pointy stone flakes onto wooden shafts. In addition to improving tool making methods, the Middle Paleolithic also saw an improvement of the tools themselves that allowed access to a wider variety and amount of food sources. For example microliths or small stone tools or points were invented around 70,000 or 65,000 BP and were essential to the invention of bows and spear throwers in the following Upper Paleolithic period.^[25] Harpoons were invented and used for the first time during the late Middle Paleolithic (c.90,000 years ago); the invention of these devices brought fish into the human diets, which provided a hedge against starvation and a more abundant food supply.^[30][31] Thanks to their technology and their advanced social structures, Paleolithic groups such as the Neanderthals who had a Middle Paleolithic level of technology, appear to have hunted large game just as well as Upper Paleolithic modern humans^[32] and the Neanderthals in particular may have likewise hunted with projectile weapons.^[33] Nonetheless, Neanderthal use of projectile weapons in hunting occurred very rarely (or perhaps never) and the Neanderthals hunted large game animals mostly by ambushing them and attacking them with mêlée weapons such as thrusting spears rather than attacking them from a distance with projectile weapons.^[20][34]

During the Upper Paleolithic, further inventions were made, such as the net (c. 22,000 or 29,000 BP)^[25] bolas,^[35] the spear thrower (c.30,000 BP), the bow and arrow (c. 25,000 or 30,000 BP)^[2][36] and the oldest example of ceramic art, the Venus of Dolní Věstonice (c. 29,000–25,000 BCE).^[2] Early dogs were domesticated, sometime between 30,000 BP and 14,000 BP, presumably to aid in hunting.^[37] However, the earliest instances of successful domestication of dogs may be much more ancient than this. Evidence from canine DNA collected by Robert K. Wayne suggests that dogs may have been first domesticated in the late Middle Paleolithic around 100,000 BP or perhaps even earlier.^[38] Archeological evidence from the Dordogne region of France demonstrates that members of the European early Upper Paleolithic culture known as the Aurignacian used calendars (c. 30,000 BP). This was a lunar calendar that was used to document the phases of the moon. Genuine solar calendars did not appear until the following Neolithic period.^[39] It is almost certain that Upper Paleolithic cultures could precisely time the migration of game animals such as wild horses and deer.^[31] This ability allowed humans to become efficient hunters and to exploit a wide variety of game animals.^[31] Recent research indicates that the Neanderthals timed their hunts and the migrations of game animals long before the beginning of the Upper Paleolithic.^[32]

Social organization[link]

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Humans may have taken part in long-distance trade between bands for rare commodities and raw materials (such as stone needed for making tools) as early as 120,000 years ago in Middle Paleolithic.

The social organization of the earliest Paleolithic (Lower Paleolithic) societies remains largely unknown to scientists, though Lower Paleolithic hominids such as Homo habilis and Homo erectus are likely to have had more complex social structures than chimpanzee societies.^[40] Late Oldowan/Early Acheulean humans such as Homo ergaster/Homo erectus may have been the first people to invent central campsites or home bases and incorporate them into their foraging and hunting strategies like contemporary hunter-gatherers, possibly as early as 1.7 million years ago;^[3] however, the earliest solid evidence for the existence of home bases or central campsites (hearths and shelters) among humans only dates back to 500,000 years ago.^[3]

Similarly, scientists disagree whether Lower Paleolithic humans were largely monogamous or polygamous.^[40] In particular, the Provisional model suggests that bipedalism arose in Pre Paleolithic australopithecine societies as an adaptation to monogamous lifestyles; however, other researchers note that sexual dimorphism is more pronounced in Lower Paleolithic humans such as Homo erectus than in Modern humans, who are less polygamous than other primates, which suggests that Lower Paleolithic humans had a largely polygamous lifestyle, because species that have the most pronounced sexual dimorphism tend more likely to be polygamous.^[41]

Human societies from the Paleolithic to the early Neolithic farming tribes lived without states and organized governments. For most of the Lower Paleolithic, human societies were possibly more hierarchical than their Middle and Upper Paleolithic descendants, and probably were not grouped into bands,^[42] though during the end of the Lower Paleolithic, the latest populations of the hominid Homo erectus may have begun living in small-scale (possibly egalitarian) bands similar to both Middle and Upper Paleolithic societies and modern hunter-gatherers.^[42]

Middle Paleolithic societies, unlike Lower Paleolithic and early Neolithic ones, consisted of bands that ranged from 20 to 30 or 25 to 100 members and were usually nomadic.^[2][42][43] These bands were formed by several families. Bands sometimes joined together into larger "macrobands" for activities such as acquiring mates and celebrations or where resources were abundant.^[2] By the end of the Paleolithic era, about 10,000 BP people began to settle down into permanent locations, and began to rely on agriculture for sustenance in many locations. Much evidence exists that humans took part in long-distance trade between bands for rare commodities (such as ochre, which was often used for religious purposes such as ritual^[44][45]) and raw materials, as early as 120,000 years ago in Middle Paleolithic.^[20] Inter-band trade may have appeared during the Middle Paleolithic because trade between bands would have helped ensure their survival by allowing them to exchange resources and commodities such as raw materials during times of relative scarcity (i.e. famine, drought).^[20] Like in modern hunter-gatherer societies, individuals in Paleolithic societies may have been subordinate to the band as a whole.^[16][17] Both Neanderthals and modern humans took care of the elderly members of their societies during the Middle and Upper Paleolithic.^[20]