Tapir Temporal range: Early Eocene–Recent PreЄ Є O S D C P T J K Pg N
Brazilian tapir
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Perissodactyla
Superfamily:	Tapiroidea
Family:	Tapiridae Gray, 1821
Genus:	Tapirus Brünnich, 1772
Species
Tapirus bairdii Tapirus indicus Tapirus pinchaque Tapirus terrestris

A tapir ( /ˈteɪpər/ TAY-pər or /təˈpɪər/ tə-PEER) is a large browsing mammal, similar in shape to a pig, with a short, prehensile snout. Tapirs inhabit jungle and forest regions of South America, Central America, and Southeast Asia. The four species of tapirs are: the Brazilian tapir, the Malayan tapir, Baird's tapir and the Mountain tapir. All four are classified as endangered or vulnerable. Their closest relatives are the other odd-toed ungulates, including horses and rhinoceroses.

Species[link]

Four extant species are widely recognized, though some authors describe more, and a number are extinct:

• Baird's tapir, Tapirus bairdii
• Brazilian tapir (also called the South American or lowland tapir), Tapirus terrestris
• Malayan tapir, Tapirus indicus
• Mountain tapir, Tapirus pinchaque
• Tapirus californicus †
• Tapirus copei †
• Tapirus merriami †
• Tapirus polkensis †
• Tapirus veroensis †

Hybrids[link]

Hybrids of the Baird's and the Brazilian tapirs were bred at the San Francisco Zoo around 1969 and produced a second generation around 1970.^[1]

A Baird's tapir in Belize

General appearance[link]

Size varies between types, but most tapirs are about 2 metres (7 ft) long, stand about a metre (3 ft) high at the shoulder, and weigh between 150 and 300 kg (330 to 700 lb). Coats are short and range in color from reddish-brown to grey to nearly black, with the notable exceptions of the Malayan tapir, which has a white, saddle-shaped marking on its back, and the mountain tapir, which has longer, woolly fur. All tapirs have oval, white-tipped ears, rounded, protruding rumps with stubby tails, and splayed, hooved toes, with four toes on the front feet and three on the hind feet, which help them to walk on muddy and soft ground. Baby tapirs of all types have striped-and-spotted coats for camouflage. Females have a single pair of mammary glands.^[2]

Physical characteristics[link]

Flehmen response

The proboscis of the tapir is a highly flexible structure, able to move in all directions, allowing the animals to grab foliage that would otherwise be out of reach. Tapirs often exhibit the flehmen response, a posture in which they raise their snouts and show their teeth to detect scents. This response is frequently exhibited by bulls sniffing for signs of other males or females in oestrus in the area. The length of the proboscis varies among species; Malayan tapirs have the longest snouts and Brazilian tapirs have the shortest.^[3] The evolution of tapir proboscises, made up almost entirely of soft tissues rather than bony internal structures, gives the Tapiridae skull a unique form in comparison to other perissodactyls, with a larger sagittal crest, orbits positioned more rostrally, a posteriorly telescoped cranium, and a more elongated and retracted nasoincisive incisure.^[4][5]

Sketch of the skull of a tapir, compared with a Malayan tapir shown in profile

Tapirs have brachyodont, or low-crowned, teeth that lack cement. Their dental formula is

totalling 42 to 44 teeth; this dentition is closer to that of equids, who may differ by one less canine, than their other perissodactyl relatives, rhinoceroses.^[6][7] Their incisors are chisel-shaped, with the third large, conical upper incisor separated by a short gap from the considerably smaller canine. A much longer gap is found between the canines and premolars, the first of which may be absent.^[8] Tapirs are lophodonts, and their cheek teeth have distinct lophs (ridges) between protocones, paracones, metacones and hypocones.^[9][10]

Tapirs have brown eyes, often with a bluish cast to them which has been identified as corneal cloudiness, a condition most commonly found in Malayan tapirs. The exact etiology is unknown, but the cloudiness may be caused by excessive exposure to light or by trauma.^[11][12] However, the tapir's sensitive ears and strong sense of smell help to compensate for deficiencies in vision.

Tapirs have simple stomachs, and, like horses, ferment food in a large cecum.^[13]

Lifecycle[link]

Young tapirs reach sexual maturity between three and five years of age, with females maturing earlier than males.^[14] Under good conditions, a healthy female tapir can reproduce every two years; a single youngster is born after a gestation of about 13 months. The natural lifespan of a tapir is approximately 25 to 30 years, both in the wild and in zoos.^[15] Apart from mothers and their young offspring, tapirs lead almost exclusively solitary lives.

Behavior[link]

The undersides of the front (left, with four toes) and back (right, with three toes) feet of a Malayan tapir at rest

Although they frequently live in dryland forests, tapirs with access to rivers spend a good deal of time in and under the water, feeding on soft vegetation, taking refuge from predators, and cooling off during hot periods. Tapirs near a water source will swim, sink to the bottom and walk along the riverbed to feed, and have been known to submerge themselves under water to allow small fish to pick parasites off their bulky bodies.^[16] Along with fresh water lounging, tapirs often wallow in mud pits, which also helps to keep them cool and free of insects.

In the wild, the tapir’s diet consists of fruit, berries, and leaves, particularly young, tender growth. Tapirs will spend many of their waking hours foraging along well-worn trails, snouts to the ground in search of food. Baird’s tapirs have been observed to eat around 40 kilograms (85 pounds) of vegetation in one day.^[17]

Tapirs are largely nocturnal and crepuscular, although the smaller Mountain tapir of the Andes is generally more active during the day than its congeners. They have monocular vision.

Copulation may occur in or out of water, and in captivity, mating pairs will often copulate multiple times during oestrus.^[18][19]

An adult Malayan tapir at the San Diego Zoo

Tooth from the extinct Tapirus veroensis 2.5cm wide. ~1 million years old, Alluvial deposits, Florida, USA

A baby Brazilian tapir with spots and stripes characteristic of all juvenile tapirs

An adult Malayan tapir sitting

Habitat, predation, and vulnerability[link]

Adult tapirs are large enough that they have few natural predators, and the thick skin on the backs of their necks helps to protect them from threats such as jaguars, crocodiles, anacondas, and tigers. The creatures are also able to run fairly quickly, considering their size and cumbersome appearance, finding shelter in the thick undergrowth of the forest or in water. Hunting for meat and hides has substantially reduced their numbers and, more recently, habitat loss has resulted in the conservation watch-listing of all four species: both the Brazilian tapir and the Malayan tapir are classified as vulnerable; and the Baird’s tapir and the Mountain tapir are endangered.

Evolution and natural history[link]

The first tapirids, such as Heptodon, appeared in the early Eocene.^[20] They appeared very similar to modern forms, but were about half the size, and lack the proboscis. The first true tapirs appeared in the Oligocene, and by the Miocene, such genera as Miotapirus were almost indistinguishable from the extant species. It is believed that Asian and American tapirs diverged around 20 to 30 million years ago, and that tapirs migrated from North America to South America around 3 million years ago, as part of the Great American Interchange.^[21] For much of their history, tapirs were spread across the northern hemisphere, where they became extinct as recently as 10,000 years ago.^[22] T. merriami, T. veroensis, T. copei, and T. californicus became extinct during the Pleistocene in North America. The giant tapir Megatapirus survived until about 4,000 years ago in China.

It is also believed by some scientists that the tapir may have evolved from the paleothere Hyracotherium (once thought to be a primitive horse).^[23]

Genetics[link]

The species of tapir have the following chromosomal numbers:

The Malayan tapir, the species most isolated geographically from the rest of the genus, has a significantly smaller number of chromosomes and has been found to share fewer homologies with the three types of American tapirs. A number of conserved autosomes (13 between karyotypes of the Baird’s tapir and Brazilian tapir, and 15 between the Baird’s and Mountain tapir) have also been found in the American species that are not found in the Asian animal. However, geographic proximity is not an absolute predictor of genetic similarity; for instance, G-banded preparations have revealed that Malayan, Baird’s and Brazilian tapirs have identical X chromosomes, while Mountain tapirs are separated by a heterochromatic addition/deletion.^[24]

Lack of genetic diversity in tapir populations has become a major source of concern for conservationists. Habitat loss has isolated already small populations of wild tapirs, putting each group in greater danger of dying out completely. Even in zoos, genetic diversity is limited; all captive mountain tapirs, for example, are descended from only two founder individuals.^[25]

Conservation[link]

There are a number of conservation projects around the world. The Tapir Specialist Group, a unit of the IUCN Species Survival Commission, strives to conserve biological diversity by stimulating, developing, and executing practical programs to study, save, restore, and manage the four species of tapir and their remaining habitats in Central and South America and Southeast Asia.^[26]

The Baird's Tapir Project of Costa Rica is the longest ongoing tapir project in the world, having started in 1994. It involves placing radio collars on tapirs in Costa Rica's Corcovado National Park to study their social systems and habitat preferences.^[27]

27 April 2008, was World Tapir Day. The day has been established to raise awareness about the four species of tapir that inhabit Central and South America and South-East Asia.^[28]

Attacks on humans[link]

A Mountain tapir, the most threatened of all the tapirs, and also the woolliest

Tapirs are generally shy, but when they are scared they can defend themselves with their very powerful jaws. In 1998, a zookeeper in Oklahoma City was mauled and had an arm severed after opening the door to a female tapir's enclosure to push food inside. (The tapir's two-month-old baby also occupied the cage at the time.)^[29] In 2006, Carlos Manuel Rodriguez Echandi (who was then the Costa Rican Environmental Minister) became lost in the Corcovado National Park and was found by a search party with a "nasty bite" from a wild tapir.^[30] However, such examples are rare; for the most part, tapirs are likely to avoid confrontation in favour of running from predators, hiding, or, if possible, submerging themselves in nearby water until a threat is gone.^[31]

Frank Buck wrote about an attack by a tapir in 1926, which he described in his book, Bring 'Em Back Alive.^[32]

Cultural references[link]

Lowland tapir earthenware from Suriname, made before 1914

In Chinese, Korean and Japanese, the tapir is named after a beast from Chinese mythology. A feature of this mythical creature is a snout like that of an elephant. In Japanese folklore, tapirs can eat people's dreams. In Chinese, the name of this beast, subsequently the name of the tapir, is mò in Mandarin (貘) and mahk in Cantonese (貘). The Korean equivalent is maek (Hangul: 맥, Hanja: 貘 [출처] 테이퍼 [貘, tapir ] ), while it is called baku (バク) in Japanese. The Chinese file hosting service Mofile has been referred to as the tapir by Chinese-speaking users.

In the prehistoric sequences of the science fiction film 2001: A Space Odyssey, tapirs appear alongside primitive hominids. The tapir was chosen because of its unusual appearance, which is frequently described as being prehistoric. However, there is no fossil-based evidence of tapirs ever existing in Africa.^[33]

Near the start of the film Ice Age (2002), a family of tapirs appear in a herd of many species, several being prehistoric. The father tapir scolds his children for playing in tar.

References[link]

External links[link]

• IUCN/SSC Tapir Specialist Group
• The Tapir Gallery at The Tapir Preservation Fund website
• World Tapir Day website
• Baird's Tapir Project of Costa Rica

Henry Doorly (November 9, 1879 – June 21, 1961), a native of Barbados, moved to Omaha while working as an engineer and then moved into the newspaper business. He became the chairman of the World Publishing Company and publisher of the Omaha World-Herald in Omaha, Nebraska. Doorly worked for the company for 58 years,^[1] becoming a highly influential figure in the city. Omaha's zoo was renamed in his memory in 1963.^[2]

Early years[link]

Born to Martin E. Doorly and Katherine Carrington in Barbados, Henry was educated at Harrison College in Bridgetown. From 1896 to 1898 he studied civil engineering in the West Indies. Arriving in Omaha in 1902 as a surveyor with the Union Pacific Railroad, he spent two years working as a draftsman with the United States Army Corps of Engineers in Omaha. On September 7, 1904 Doorly married Margaret Hitchcock in Omaha, becoming the son-in-law of World-Herald publisher and politician Gilbert M. Hitchcock.^[3]

Omaha World-Herald[link]

Beginning as a reporter for the Omaha World-Herald, Doorly failed miserably, retaining his job only because he was the publisher's daughter's fiancé. Doorly became successful after moving to advertisement sales, soon becoming advertising manager and then business manager for the newspaper.^[4]

Doorly took control of the Omaha World-Herald in 1934 when his father-in-law, Gilbert M. Hitchcock, died. Reflecting the changing nature of the major American political parties and Doorly's personal disenchantment with the New Deal in the 1930s, he implemented the newspaper's editorial page shift toward a Republican Party policy stance.

Under Doorly's guidance, the paper soon standardized advertisement policies and procedures. To enforce brevity and variety, Doorly had a daily "Item Count" conducted to count the number of stories in each news category, including local news, society, and international sections. The staff consequently produced as many as 450 separate news stories a day.^[5]

Omaha Bee[link]

In 1937 William Randolph Hearst sold Doorly the Bee-News, his main competitor in Omaha.^[6] During World War II, Doorly used his position to lead a national campaign educating newspaper editors and publishers in promoting steel recycling to support the war.^[7]

Doorly retired in 1955, leaving control of both newspapers to Walter E. Christiansen.^[5] He died in 1961 of an apparent heart attack.^[8][9]

Legacy[link]

In 1963, his widow Margaret Hitchcock Doorly donated $750,000 (approximately $4.5 million in 2005 dollars) to the Omaha Zoological Society. It was organized in 1953 to improve the Riverview Park Zoo and to provide administrative help to the city.^[10] With her donation, Doorly stipulated that the zoo be renamed in memory of her late husband.^[2]

See also[link]

• History of Omaha

References[link]