The giant panda, or panda (''Ailuropoda melanoleuca'', literally meaning "black and white cat-foot") is a bear native to central-western and south western China. It is easily recognized by its large, distinctive black patches around the eyes, over the ears, and across its round body. Though it belongs to the order Carnivora, the panda's diet is 99% bamboo. Pandas in the wild will occasionally eat other grasses, wild tubers, or even meat in the form of birds, rodents or carrion. In captivity they may receive honey, eggs, fish, yams, shrub leaves, oranges, or bananas along with specially prepared feed.
The giant panda lives in a few mountain ranges in central China, mainly in Sichuan province, but also in the Shaanxi and Gansu provinces. Due to farming, deforestation and other development, the panda has been driven out of the lowland areas where it once lived.
The panda is a conservation reliant endangered species. A 2007 report shows 239 pandas living in captivity inside China and another 27 outside the country. Wild population estimates vary; one estimate shows that there are about 1,590 individuals living in the wild, while a 2006 study via DNA analysis estimated that this figure could be as high as 2,000 to 3,000. Some reports also show that the number of pandas in the wild is on the rise. However, the IUCN does not believe there is enough certainty yet to reclassify the species from Endangered to Vulnerable.
While the dragon has historically served as China's national emblem, in recent decades the panda has also served as an emblem for the country. Its image appears on a large number of modern Chinese commemorative silver, gold, and platinum coins. Though the panda is often assumed to be docile, it has been known to attack humans, presumably out of irritation rather than predation.
The giant panda has a body shape typical of bears. It has black fur on its ears, eye patches, muzzle, legs, arms and shoulders. The rest of the animal's coat is white. Although scientists do not know why these unusual bears are black and white, some speculate that the bold coloring provides effective camouflage in its shade-dappled snowy and rocky surroundings. The giant panda's thick, wooly coat keeps it warm in the cool forests of its habitat. The giant panda has large molar teeth and strong jaw muscles for crushing tough bamboo.
The giant panda's paw has a "thumb" and five fingers; the "thumb" is actually a modified sesamoid bone, which helps the giant panda to hold bamboo while eating. Stephen Jay Gould discusses this feature in his book of essays on evolution and biology, ''The Panda's Thumb''.
The giant panda's tail, measuring , is the second longest in the bear family. The longest belongs to the Sloth Bear.
The giant panda typically lives around 20 years in the wild and up to 30 years in captivity. The recorded age of the oldest captive giant panda, a female named Ming Ming, is 34.
Social encounters occur primarily during the brief breeding season in which pandas in proximity to one another will gather. After mating, the male leaves the female alone to raise the cub.
Despite its taxonomic classification as a carnivoran, the giant panda's diet is primarily herbivorous, consisting almost exclusively of bamboo. However, the giant panda still has the digestive system of a carnivore, as well as carnivore-specific genes, and thus derives little energy and little protein from consumption of bamboo. Its ability to digest cellulose is ascribed to the microbes in its gut. The giant panda is a "highly specialized" animal with "unique adaptations", and has lived in bamboo forests for millions of years. The average giant panda eats as much as 9 to 14 kg (20 to 30 pounds) of bamboo shoots a day. Because the giant panda consumes a diet low in nutrition, it is important for it to keep its digestive tract full. The limited energy input imposed on it by its diet has affected the panda's behavior. The giant panda tends to limit its social interactions and avoids steeply sloping terrain in order to limit its energy expenditures.
Two of the panda's most distinctive features, its large size and its round face, are adaptations to its bamboo diet. Panda researcher Russell Ciochon observed that: “[much] like the vegetarian gorilla, the low body surface area to body volume [of the giant panda] is indicative of a lower metabolic rate. This lower metabolic rate and a more sedentary lifestyle allow the giant panda to subsist on nutrient poor resources such as bamboo.” Similarly, the giant panda's round face is the result of powerful jaw muscles, which attach from the top of the head to the jaw. Large molars crush and grind fibrous plant material.
Pandas eat any of twenty-five bamboo species in the wild, such as ''Fargesia dracocephala'' and ''Fargesia rufa''. Only a few bamboo species are widespread at the high altitudes pandas now inhabit. Bamboo leaves contain the highest protein levels; stems have less. Given this large diet, the giant panda can defecate up to 40 times a day.
Because of the synchronous flowering, death, and regeneration of all bamboo within a species, the giant panda must have at least two different species available in its range to avoid starvation. While primarily herbivorous, the giant panda still retains decidedly ursine teeth, and will eat meat, fish, and eggs when available. In captivity, zoos typically maintain the giant panda's bamboo diet, though some will provide specially-formulated biscuits or other dietary supplements.
Despite the shared name, habitat type, and diet, as well as a unique enlarged bone called the ''pseudo thumb'' (which helps them grip the bamboo shoots they eat), the giant panda and red panda are only distantly related. Molecular studies have placed the red panda in its own family Ailuridae, and not under Ursidae.
The creature named ''mo'' (貘) mentioned in some ancient books has been interpreted as giant panda. The dictionary ''Shuowen Jiezi'' (Eastern Han Dynasty) says that the ''mo'', from Shu (Sichuan), is bear-like, but yellow-and-black, although the older ''Erya'' describes ''mo'' simply as a "white leopard". The interpretation of the legendary fierce creature ''pixiu'' (貔貅) as referring to the giant panda are also common.
During the reign of the Yongle Emperor (early 15th century), his relative from Kaifeng sent him a captured ''zouyu'' (騶虞), and another ''zouyu'' was sighted in Shandong. ''Zouyu'' is a legendary "righteous" animal, which, similarly to a ''qilin'', only appears during the rule of a benevolent and sincere monarch. It is said to be fierce as a tiger, but gentle and strictly vegetarian, and described in some books as a white tiger with black spots. Puzzled about the real zoological identity of the creature captured during the Yongle era, J.J.L. Duyvendak exclaims, "Can it possibly have been a Pandah?"
The comparative obscurity of the giant panda throughout most of China's history is illustrated by the fact that, despite there being a number of depictions of bears in Chinese art starting from its most ancient times, and the bamboo being one of the favorite subjects for Chinese painters, there are no known pre-20th-century artistic representations of giant pandas.
By 1984, however, pandas were no longer used as agents of diplomacy. Instead, the PRC began to offer pandas to other nations only on 10-year loans. The standard loan terms include a fee of up to US$1,000,000 per year and a provision that any cubs born during the loan are the property of the People's Republic of China. Since 1998, due to a WWF lawsuit, the United States Fish and Wildlife Service only allows a U.S. zoo to import a panda if the zoo can ensure that the PRC will channel more than half of its loan fee into conservation efforts for the giant panda and its habitat.
In May 2005, the People's Republic of China offered a breeding pair to Taiwan. The issue became embroiled in cross-Strait relations—both over the underlying symbolism, and over technical issues such as whether the transfer would be considered "domestic" or "international," or whether any true conservation purpose would be served by the exchange. A contest in 2006 to name the pandas was held in the mainland, resulting in the politically charged names Tuan Tuan and Yuan Yuan (from ''tuanyuan'', meaning "reunion", i.e. "reunification"). PRC's offer was initially rejected by President Chen of Taiwan. However when Ma Ying-jeou assumed the presidency in 2008 the offer was accepted, and the pandas arrived in December of that year.
The giant panda has been a target for poaching by locals since ancient times and by foreigners since it was introduced to the West. Starting in the 1930s, foreigners were unable to poach giant pandas in China because of the Second Sino-Japanese War and the Chinese Civil War, but pandas remained a source of soft furs for the locals. The population boom in China after 1949 created stress on the pandas' habitat, and the subsequent famines led to the increased hunting of wildlife, including pandas. During the Cultural Revolution, all studies and conservation activities on the pandas were stopped. After the Chinese economic reform, demand for panda skins from Hong Kong and Japan led to illegal poaching for the black market, acts generally ignored by the local officials at the time.
Though the Wolong National Nature Reserve was set up by the PRC government in 1958 to save the declining panda population, few advances in the conservation of pandas were made, due to inexperience and insufficient knowledge of ecology. Many believed that the best way to save the pandas was to cage them. As a result, pandas were caged at any sign of decline, and suffered from terrible conditions. Because of pollution and destruction of their natural habitat, along with segregation due to caging, reproduction of wild pandas was severely limited. In the 1990s, however, several laws (including gun control and the removal of resident humans from the reserves) helped the chances of survival for pandas. With these renewed efforts and improved conservation methods, wild pandas have started to increase in numbers in some areas, even though they still are classified as a rare species.
In 2006, scientists reported that the number of pandas living in the wild may have been underestimated at about 1,000. Previous population surveys had used conventional methods to estimate the size of the wild panda population, but using a new method that analyzes DNA from panda droppings, scientists believe that the wild panda population may be as large as 3,000. Although the species is still endangered, it is thought that the conservation efforts are working. In 2006, there were 40 panda reserves in China, compared to just 13 reserves two decades ago.
The giant panda is among the world's most adored and protected rare animals, and is one of the few in the world whose natural inhabitant status was able to gain a UNESCO World Heritage Site designation. The Sichuan Giant Panda Sanctuaries, located in the southwest Sichuan province and covering seven natural reserves, were inscribed onto the World Heritage List in 2006.
Not all conservationists agree that the money spent on conserving pandas is money well spent. Chris Packham has argued that breeding pandas in captivity is "pointless" because "there is not enough habitat left to sustain them". Packham argues that the money spent on pandas would be better spent elsewhere, and has said that he would "eat the last panda if I could have all the money we have spent on panda conservation put back on the table for me to do more sensible things with," though he has apologized for upsetting people who like pandas. He points out that "The panda is possibly one of the grossest wastes of conservation money in the last half century."
Initially the primary method of breeding giant pandas in captivity was by artificial insemination, as they seemed to lose their interest in mating once they were captured. This led some scientists to try extreme methods such as showing them videos of giant pandas mating and giving the males Viagra. Only recently have researchers started having success with captive breeding programs, and they have now determined that giant pandas have comparable breeding to some populations of the American black bear, a thriving bear family. The current reproductive rate is considered one young every two years.
Giant pandas reach sexual maturity between the ages of four and eight, and may be reproductive until age 20. The mating season is between March and May, when a female goes into her estrous cycle which lasts for two or three days and only occurs once a year. When mating, the female is in a crouching, head-down position as the male mounts her from behind. Copulation time is short, ranging from thirty seconds to five minutes, but the male may mount her repeatedly to ensure successful fertilization. The gestation period ranges from 95 to 160 days. Cubs weigh only 90 to 130 grams (3.2 to 4.6 ounces), which is about 1/800 of the mother's weight.
If twins are born, usually only one survives in the wild. The mother will select the stronger of the cubs, and the weaker will die. It is thought that the mother cannot produce enough milk for two cubs since she does not store fat. The father has no part in helping raise the cub.When the cub is first born, it is pink, blind, and toothless. A giant panda cub is also extremely small, and it is difficult for the mother to protect it because of the baby's size. It nurses from its mother's breast 6 to 14 times a day for up to 30 minutes at a time. For three to four hours, the mother may leave the den to feed, which leaves the cub defenseless. One to two weeks after birth, the cub's skin turns gray where its hair will eventually become black. A slight pink color may appear on cub's fur, as a result of a chemical reaction between the fur and its mother's saliva. A month after birth, the color pattern of the cub's fur is fully developed. A cub's fur is very soft and coarsens with age. The cub begins to crawl at 75 to 80 days; mothers play with their cubs by rolling and wrestling with them. The cubs are able to eat small quantities of bamboo after six months, though mother's milk remains the primary food source for most of the first year. Giant panda cubs weigh 45 kg (100 pounds) at one year, and live with their mothers until they are 18 months to two years old. The interval between births in the wild is generally two years.
In July 2009, Chinese scientists confirmed the birth of the first cub to be successfully conceived through artificial insemination using frozen sperm. The cub was born at 07:41 on 23 July that year in Sichuan as the third cub of You You, an 11-year-old. The technique for freezing the sperm in liquid nitrogen was first developed in 1980 and the first birth was hailed as a solution to the problem of lessening giant panda semen availability which had led to in-breeding. It has been suggested that panda semen, which can be frozen for decades, could be shared between different zoos to save the species. It is expected that zoos in destinations such as San Diego in the United States and Mexico City will now be able to provide their own semen to inseminate more giant pandas.
Attempts have also been made to reproduce giant pandas by interspecific pregnancy by implanting cloned panda embryos into the uterus of an animal of another species. This has resulted in panda fetuses, but no live births.
In most encyclopedic sources, the name "panda" or "common panda" originally referred to the lesser-known red panda, thus necessitating the inclusion of "giant" and "lesser/red" prefixes in front of the names. Even in 2010, the ''Encyclopædia Britannica'' still used "giant panda" or "panda bear" for the bear and simply "panda" for the Ailuridae, despite the popular usage of the word "panda".
Since the earliest collection of Chinese writings, the Chinese language has given the bear 20 different names, such as 花熊 (hua xiong) "spotted bear" and 竹熊 (zhu xiong) "bamboo bear". The most popular names in China today are 大熊貓 (dà xióng māo), literally "large bear cat", or just 熊貓 (xióng māo), "bear cat". The name may have been inspired by the giant panda's eyes, which have pupils that are cat-like vertical slits – unlike other bear species, which have round pupils.
In Taiwan, the popular name for panda is the inverted 貓熊 (māo xióng) "cat bear," even though many encyclopedia and dictionaries in Taiwan still use "bear cat" as the correct name. Some linguists argue that, in this construction, "bear" instead of "cat" is the base noun, making this name more grammatically and logically correct, which may have led to the popular choice despite official writings.
Pandas have been kept in zoos as early as the Western Han Dynasty in China, where the writer Sima Xiangru notes that the panda was the most treasured animal in the emperor's garden of exotic animals in the capital Chang'an (present Xi'an). Not until the 1950s were pandas again recorded to have been exhibited in China's zoos.
Chi Chi at the London Zoo became very popular. This influenced the World Wildlife Fund to use a panda as its symbol.
A 2006 ''New York Times'' article outlined the economics of keeping pandas, which costs five times more than that of the next most expensive animal, an elephant. American zoos generally pay the Chinese government $1 million a year in fees, as part of a typical ten-year contract. San Diego's contract with China was to expire in 2008 but got a five-year extension at about half of the previous yearly cost. The last contract, with the Memphis Zoo in Memphis, Tennessee, ends in 2013.
;Other places in Asia Taipei Zoo, Taipei, Taiwan – home to Tuan Tuan (M) and Yuan Yuan (F). Chiang Mai Zoo, Chiang Mai, Thailand – home to Chuang Chuang (M), Lin Hui (F), and Lin Bing, a female cub born 27 May 2009 Adventure World, Shirahama, Wakayama – Until recently, home to Ei Mei (M), Mei Mei (F), Rau Hin (F), Ryu Hin and Syu Hin (male twins), and Kou Hin (M). In December 2006, twin cubs were born to Ei Mei and Mei Mei. Two cubs, Eiihin (M) and Meihin (F), were born to Rau Hin on 13 September 2008. Mei Mei, a mother of ten cubs, died on 15 October 2008. Oji Zoo, Kobe, Hyōgo – home of Kou Kou (M), Tan Tan (F) River Safari, a new park under Wildlife Reserves Singapore, Singapore – to receive two pandas (Kai Kai and Jia Jia) in 2012.
Recently, Natural History New Zealand (NHNZ) has featured pandas in two documentaries. ''Panda Nursery'' (2006) featured China’s Wolong National Nature Reserve in the mountains in Sichuan Province; forty giant pandas and a dedicated team of staff play a crucial role in ensuring the survival of the species. As part of the Reserve’s panda breeding program, a revolutionary new method of rearing twin cubs called ‘swap-raising’ has been developed. Each cub is raised by both its natural mother and one of the Reserve’s veterinarians, Wei Rongping, to increase the chances of both cubs surviving. ''Growing Up: Giant Panda'' (2003) featured Chengdu Giant Panda Center in south-west China as one of the best in the world. Yet with female pandas' short fertility cycles and low birth rates, raising the captive panda population is an uphill battle.
;Bibliography
Category:Giant pandas Category:National symbols of China Category:EDGE Species Category:Conservation reliant species Category:Articles containing video clips Category:Megafauna of Eurasia Category:Mammals of China Category:Herbivorous animals Category:Sequenced genomes Category:Animals described in 1869
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