{{automatic taxobox | name=Amphibians | fossil_range= | image = Oophaga.pumilio.zoo.jpg | image_caption = Strawberry Poison-dart Frog, '' Oophaga pumilio'' | taxon = Amphibia | authority = Linnaeus, 1758 | subdivision_ranks = Subclasses and Orders | subdivision =    Order Temnospondyli – ''extinct'' Subclass Lepospondyli – ''extinct'' Subclass Lissamphibia    Order Anura    Order Caudata    Order Gymnophiona}}

Amphibians (class Amphibia, from Amphi- meaning "on both sides" and -bios meaning "life"), are a class of vertebrate animals including animals such as frogs, caecilians, and salamanders. They are characterized as non-amniote ectothermic (or cold-blooded) tetrapods. Most Amphibians undergo metamorphosis from a juvenile water-breathing form to an adult air-breathing form, but some are paedomorphs that retain the juvenile water-breathing form throughout life. Mudpuppies, for example, retain juvenile gills in adulthood. The three modern orders of amphibians are Anura (frogs and toads), Caudata (salamanders and newts), and Gymnophiona (caecilians, limbless amphibians that resemble snakes), and in total they number approximately 6,500 species. Many amphibians lay their eggs in water. Amphibians are superficially similar to reptiles, but reptiles are amniotes, along with mammals and birds. The study of amphibians is called batrachology.

Amphibians are ecological indicators, and in recent decades there has been a dramatic decline in amphibian populations around the globe. Many species are now threatened or extinct.

Amphibians evolved in the Devonian Period and were top predators in the Carboniferous and Permian Periods, but many lineages were wiped out during the Permian–Triassic extinction. One group, the metoposaurs, remained important predators during the Triassic, but as the world became drier during the Early Jurassic they died out, leaving a handful of relict temnospondyls like ''Koolasuchus'' and the modern orders of Lissamphibia.

Etymology

''Amphibian'' is derived from the Ancient Greek term ἀμφίβιος ''amphíbios'' which means both kinds of life, ''amphi'' meaning “both” and ''bio'' meaning life. The term was initially used for all kinds of combined natures. Eventually it was used to refer to animals that live both in the water and on land.

Evolutionary history

The first major groups of amphibians developed in the Devonian Period from fish similar to the modern coelacanth and lungfish which had evolved multi-jointed leg-like fins that enabled them to crawl along the sea bottom. These amphibians were as much as one to five meters in length. However, amphibians never developed the ability to live their entire lives on land, having to return to water to lay their shell-less eggs.

In the Carboniferous Period, the amphibians moved up in the food chain and began to occupy the ecological position currently occupied by crocodiles. These amphibians were notable for eating the mega insects on land and many types of fishes in the water. During the Triassic Period, the better land-adapted proto-crocodiles began to compete with amphibians, leading to their reduction in size and importance in the biosphere.

Taxonomic history

Traditionally, amphibians have included all tetrapod vertebrates that are not amniotes. They are divided into three subclasses, of which two are only known as extinct subclasses:

Subclass Labyrinthodontia† (diverse Paleozoic and early Mesozoic group)

Subclass Lepospondyli† (small Paleozoic group, sometimes included in the Labyrinthodontia, which may actually be more closely related to amniotes than Lissamphibia)

Subclass Lissamphibia (frogs, toads, salamanders, newts, etc.)

Of these only the last subclass includes recent species.

With the phylogenetic classification Labyrinthodontia has been discarded as it is a paraphyletic group without unique defining features apart from shared primitive characteristics. Classification varies according to the preferred phylogeny of the author, whether they use a stem-based or node-based classification. Traditionally, amphibians as a class are defined as all tetrapods with a larval stage, while the group that includes the common ancestors of all living amphibians (frogs, salamanders and caecilians) and all their descendants is called Lissamphibia. The phylogeny of Paleozoic amphibians is by no means satisfactory understood, and lissamphibia may possibly include extinct groups like the temnospondyls (traditionally placed in the subclass “Labyrinthodontia”), and the Lepospondyls, and in some analysis even the amniots. This means that phylogenetic nomenclature list a large number of basal Devonian and Carboniferous tetrapod groups, undoubtedly were “amphibians” in biology, that are formally placed in Amphibia in Linnaean taxonomy, but not in cladistic taxonomy.

All recent amphibians are included in the subclass Lissamphibia, superorder Salientia, which is usually considered a clade (which means that it is thought that they evolved from a common ancestor apart from other extinct groups), although it has also been suggested that salamanders arose separately from a temnospondyl-like ancestor, and even that caecilians are the sister group of the advanced reptiliomorph amphibians, and thus of amniots.

Authorities also disagree on whether Salientia is a Superorder that includes the order Anura, or whether Anura is a sub-order of the order Salientia. Practical considerations seem to favor using the former arrangement now. The Lissamphibia, superorder Salientia, are traditionally divided into three orders, but an extinct salamander-like family, the Albanerpetontidae, is now considered part of the Lissamphibia, besides the superorder Salientia. Furthermore, Salientia includes all three recent orders plus a single Triassic proto-frog, ''Triadobatrachus''.

Class Amphibia

Subclass Lissamphibia

** Family Albanerpetontidae — Jurassic to Miocene (extinct)

* Superorder Salientia

** Genus ''Triadobatrachus'' — Triassic (extinct) - A stem Anuran

** Order Anura (frogs and toads): Jurassic to recent — 5,602 recent species in 48 families

** Order Caudata or Urodela (salamanders, newts): Jurassic to recent — 571 recent species in 9 families

** Order Gymnophiona or Apoda (caecilians): Jurassic to recent — 174 recent species in 3 families

The actual number of species partly also depends on the taxonomic classification followed, the two most common classifications being the classification of the website AmphibiaWeb, University of California (Berkeley) and the classification by herpetologist Darrel Frost and The American Museum of Natural History, available as the online reference database Amphibian Species of the World. The numbers of species cited above follow Frost.

Respiration

The lungs in amphibians are primitive compared to that of the amniotes, possessing few internal septa, large alveoli and therefore a slow diffusion rate of oxygen into the blood. Ventilation is accomplished by buccal pumping. However, most amphibians are able to exchange gasses with the water or air via their skin. To enable sufficient cutaneous respiration, the surface of their highly vascularized skin must remain moist in order for the oxygen to diffuse at a sufficient rate. Because oxygen concentration in the water increases at both low temperatures and high flow rates, aquatic amphibians in these situations can rely primarily on cutaneous respiration, as in the Titicaca water frog and hellbender salamanders. In air, where oxygen is more concentrated, some small species can rely solely on cutaneous gas exchange, most famously the plethodontid salamanders which have neither lungs nor gills. Many aquatic salamanders and all tadpoles have gills in their larval stage, with some (such as the axolotl) retaining gills as aquatic adults.

Reproduction

For the purpose of reproduction most amphibians require fresh water. A few (e.g. ''Fejervarya raja'') can inhabit brackish water and even survive (though not thrive) in seawater, but there are no true marine amphibians. Several hundred frog species in adaptive radiations (e.g., ''Eleutherodactylus'', the Pacific Platymantines, the Australo-Papuan microhylids, and many other tropical frogs), however, do not need any water for breeding in the wild. They reproduce via direct development, an ecological and evolutionary adaptation that has allowed them to be completely independent from free-standing water. Almost all of these frogs live in wet tropical rainforests and their eggs hatch directly into miniature versions of the adult, passing through the tadpole stage within the egg. Reproductive success of many amphibians is dependent not only on the quantity of rainfall, but the seasonal timing.

Several species have also adapted to arid and semi-arid environments, but most of them still need water to lay their eggs. Symbiosis with single celled algae that lives in the jelly-like layer of the eggs has evolved several times. The larvae of frogs (tadpoles or polliwogs) breathe with exterior gills at the start, but soon a pouch is formed that covers the gills and the front legs. Lungs are also formed quite early to assist in breathing. Newt larvae have large external gills that gradually disappear and the larvae of newts are quite similar to the adult form from early age on.

Frogs and toads however have a tadpole stage, which is a totally different organism that is a grazing algae or ongrowth or filtering plankton until a certain size has been reached, where metamorphosis sets in. This metamorphosis typically lasts only 24 hours and consists of:

The disappearance of the gill pouch, making the front legs visible.

The transformation of the jaws into the big jaws of predatory frogs (most tadpoles are scraping of algae or are filter feeders)

The transformation of the digestive system: the long spiral gut of the larva is being replaced by the typical short gut of a predator.

An adaptation of the nervous system for stereoscopic vision, locomotion and feeding

A quick growth and movement of the eyes to higher up the skull and the formation of eyelids.

Formation of skin glands, thickening of the skin and loss of the lateral line system

An eardrum is developed to lock the middle ear.

The disappearance of the tail is somewhat later (occurs at higher thyroxin levels) and after the tail has been resorbed the animals are ready to leave the water. The material of the tail is being used for a quick growth of the legs. The disappearance of the larval structures is a regulated process called apoptosis.

The transformation of newts when leaving the water is reversible except for the loss of the external gills. When the animals enter the water again for reproduction changes are driven by prolactin, when they return to the land phase by thyroxin

Conservation

Dramatic declines in amphibian populations, including population crashes and mass localized extinction, have been noted in the past two decades from locations all over the world, and amphibian declines are thus perceived as one of the most critical threats to global biodiversity. A number of causes are believed to be involved, including habitat destruction and modification, over-exploitation, pollution, introduced species, climate change, endocrine-disrupting pollutants, destruction of the ozone layer (ultraviolet radiation has shown to be especially damaging to the skin, eyes, and eggs of amphibians), and diseases like chytridiomycosis. However, many of the causes of amphibian declines are still poorly understood, and are a topic of ongoing discussion. A global strategy to stem the crisis has been released in the form of the Amphibian Conservation Action Plan (available at http://www.amphibians.org). Developed by over 80 leading experts in the field, this call to action details what would be required to curtail amphibian declines and extinctions over the next 5 years - and how much this would cost. The Amphibian Specialist Group of the World Conservation Union (IUCN) is spearheading efforts to implement a comprehensive global strategy for amphibian conservation. Amphibian Ark is an organization that was formed to implement the ex-situ conservation recommendations of this plan, and they have been working with zoos and aquaria around the world encouraging them to create assurance colonies of threatened amphibians. One such project is the Panama Amphibian Rescue and Conservation Project that built on existing conservation efforts in Panama to create a country-wide response to the threat of chytridiomycosis rapidly spreading into eastern Panama

On January 21, 2008, Evolutionarily Distinct and Globally Endangered (EDGE), as given by chief Helen Meredith, identified nature's most endangered species: "The EDGE amphibians are amongst the most remarkable and unusual species on the planet and yet an alarming 85% of the top 100 are receiving little or no conservation attention." The top 10 endangered species (in the List of endangered animal species) include: the Chinese giant salamander, a distant relative of the newt, the tiny Gardiner's Seychelles, the limbless Sagalla caecilian, South African ghost frogs, lungless Mexican salamanders, the Malagasy rainbow frog, Chile's Darwin frog (Rhinoderma rufum) and the Betic Midwife Toad.

See also

Vegetal rotation

List of amphibians

References

Further reading

Solomon Berg Martin, ''Biology''

776 pages

External links

Save the Frogs!

Amphibian Specialist Group

AmphibiaWebEcuador

Amphibians photos Brazil

Amphibian Ark

AmphibiaWeb

Global Amphibian Assessment

Amphibians of central Europe

USGS--Online Guide for the Identification of Amphibians in North America north of Mexico

General amphibian biology information - Living UnderWorld

Panama Amphibian Rescue and Conservation Project

Atlanta Botanical Garden Amphibian Conservation Program

Amphibian vocalisations on Archival Sound Recordings

Froglife (British conservation charity)

Salamandridae

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name	Conan O'Brien
birth name	Conan Christopher O'Brien
alias	Coco
birth date	April 18, 1963
birth place	Brookline, Massachusetts, United States
medium	Television
nationality	American
active	1985–present
genre	Improvisational comedy, sketch comedy, physical comedy, surreal humor, self-deprecation
subjects	Self-deprecation, pop culture
influences	Johnny Carson, David Letterman, Robert Smigel
spouse	Elizabeth Ann Powel (since 2002; 2 children)
notable work	''The Simpsons''(writer, producer, 1991–1993)''Late Night with Conan O'Brien''(host, 1993–2009)''The Tonight Show with Conan O'Brien''(host, 2009–2010) ''Conan'' (host, 2010–present)
education	Harvard University
signature	Conan O'Brien Signature.svg

}} Conan Christopher O'Brien (born April 18, 1963) is an American television host, comedian, writer, producer and performer. Since November 2010 he has hosted ''Conan'', a late-night talk show that airs on the American cable television station TBS.

O'Brien was born in Brookline, Massachusetts and raised in an Irish Catholic family. He served as president of the ''Harvard Lampoon'' while attending Harvard University, and was a writer for the sketch comedy series ''Not Necessarily the News''. After writing for several comedy shows in Los Angeles, he joined the writing staff of ''Saturday Night Live'', and later of ''The Simpsons''. He hosted ''Late Night with Conan O'Brien'' from 1993 to 2009, followed by seven months hosting ''The Tonight Show with Conan O'Brien'', the only person to serve as host for both NBC programs.

Early life

O'Brien was born in Brookline, Massachusetts (a suburb of Boston) to Thomas O'Brien, a physician, epidemiologist, and professor of medicine at Harvard, and Ruth O'Brien (née Reardon), an attorney and partner at the Boston firm Ropes & Gray. He is the third of six children. O'Brien's family is Irish Catholic and descends from pre-Civil War era immigrants. In a ''Late Night'' episode, O'Brien paid a visit to County Kerry, Ireland, where his ancestors originated.

O'Brien attended Brookline High School, where he served as the managing editor of the school newspaper. In his senior year, O'Brien won the National Council of Teachers of English writing contest with his short story, "To Bury the Living". After graduating as valedictorian in 1981, he entered Harvard University. At Harvard, O'Brien lived in Holworthy Hall during his freshman year and Mather House during his three upper-class years. He concentrated in history and literature and graduated ''magna cum laude'' in 1985. His senior thesis concerned the use of children as symbols in the works of William Faulkner and Flannery O'Connor. Throughout college, O'Brien was a writer for the ''Harvard Lampoon'' humor magazine. He also briefly served as the drummer in a band called "The Bad Clams". During his sophomore and junior years, he served as the ''Lampoon'''s president. At this time, O'Brien's future boss at NBC, Jeff Zucker, was serving as President of the rival ''The Harvard Crimson''.

Career

''Saturday Night Live'' (1987–1991)

O'Brien moved to Los Angeles after graduation to join the writing staff of HBO's ''Not Necessarily the News''. He was also a writer on the short-lived ''The Wilton North Report''. He spent two years with that show and performed regularly with improvisational groups, including The Groundlings. In January 1988, ''Saturday Night Live'''s executive producer, Lorne Michaels, hired O'Brien as a writer. During his three years on ''Saturday Night Live (SNL)'', he wrote such recurring sketches as "Mr. Short-Term Memory" and "The Girl Watchers"; the latter was first performed by Tom Hanks and Jon Lovitz. O'Brien also co-wrote the sketch, "Nude Beach", with Robert Smigel, in which the word "penis" was said or sung at least 42 times. While on a writers' strike from ''Saturday Night Live'' following the 1987–88 season, O'Brien put on an improvisational comedy revue in Chicago with fellow ''SNL'' writers Bob Odenkirk and Robert Smigel called ''Happy Happy Good Show''. While living in Chicago, O'Brien briefly roomed with Jeff Garlin. In 1989, O'Brien and his fellow ''SNL'' writers received an Emmy Award for Outstanding Writing in a Comedy or Variety Series.

O'Brien, like many ''SNL'' writers, occasionally appeared as an extra in sketches; his most notable appearance was as a doorman in a sketch in which Tom Hanks was inducted into the SNL "Five-Timers Club" for hosting his fifth episode. O'Brien returned to host the show in 2001 during its 26th season. O'Brien and Robert Smigel wrote the television pilot for ''Lookwell'' starring Adam West, which aired on NBC in 1991. The pilot never went to series, but it became a cult hit. It was later screened at ''The Other Network'', a festival of unaired TV pilots produced by Un-Cabaret; it featured an extended interview with O'Brien and was rerun in 2002 on the Trio network.

''The Simpsons'' (1991–1993)

From 1991 to 1993, O'Brien was a writer and producer for ''The Simpsons'' and was credited as writer or co-writer of four episodes. Of all the episodes he wrote, he considers "Marge vs. the Monorail" to be his favorite. Along with this episode, he has sole writing credits on "New Kid on the Block," "Homer Goes to College," and "Treehouse of Horror IV," on which he wrote the episode wraparounds. He was an active producer during seasons 4 and 5 as well, meaning he would frequently contribute to scripts from those seasons as well as come up with story ideas, plot points, and jokes. The style of the show's comedy during this period was also influenced by his sensibilities, with "Marge vs. the Monorail" being cited by several former writers as the turning point in the show's history where more absurd and visual comedy became acceptable. He also developed a reputation as a "room guy," or a writer who performs comedy bits throughout the day to entertain other writers.

In his speech given at Class Day at Harvard in 2000, O'Brien credited ''The Simpsons'' with saving him, a reference to the career slump he was experiencing prior to his being hired for the show.

During his time at ''The Simpsons'', O'Brien also had a side project working with former writing partner Robert Smigel on the script for a musical film based on the "Hans and Franz" sketch from ''Saturday Night Live''. The film was never produced.

''Late Night'' (1993–2009)

As executive producer, Lorne Michaels invited O'Brien to audition to host the successor show to ''Late Night with David Letterman''. Premiering on September 13, 1993, ''Late Night with Conan O'Brien'' received generally unfavorable critical reviews during its first few years. The show remained on multiweek renewal cycles while NBC decided its fate. O'Brien later poked fun at the first three years of the show, when on his 10th Anniversary Special Mr. T appeared to give O'Brien a gold necklace with a giant "7" on it. When O'Brien tried to point out that he had actually been on the air for ten years, Mr. T responded, "I know that, fool; but you've only been funny for seven!"

Beginning in 1996, O'Brien and the ''Late Night'' writing team were nominated annually for the Emmy Award for Best Writing in a Comedy or Variety Series, winning the award for the first and only time in 2007. In 1997, 2000, 2002, 2003, and 2004, he and the ''Late Night'' writing staff won the Writers Guild Award for Best Writing in a Comedy/Variety Series. In 2001, he formed his own television production company, Conaco, which subsequently shared in the production credits for ''Late Night''.

A long-running joke, which stems from the recurring segment "Conan O'Brien Hates My Homeland", is that O'Brien resembles the first female president of Finland, Tarja Halonen. After joking about this for several months (which led to his endorsement of her campaign), O'Brien traveled to Finland, appearing on several television shows and meeting President Halonen. The trip was filmed and aired as a special.

O'Brien ad-libbed the fictional website name "hornymanatee.com" on December 4, 2006, after a sketch about the fictional manatee mascot and its inappropriate webcam site. NBC opted to purchase the website domain name for $159, since the website did not previously exist. The network was concerned that the Federal Communications Commission would hold NBC liable for promoting inappropriate content if a third party were to register the domain and post such material. For a period of time, the website hosted material concerning Conan's initial manatee joke and other ''Tonight Show'' references, but today the site just redirects to NBC's main web page.

A popular recurring bit on the show was "Pale Force", a series of animated episodes in which comedian Jim Gaffigan and O'Brien are superheroes who fight crime with their "paleness". As Gaffigan introduced each new episode, O'Brien protested the portrayal of his character as cowardly, weak, and impotent. , ''Late Night with Conan O'Brien'' had for eleven years consistently attracted an audience averaging about 2.5 million viewers. O'Brien is an avid guitarist and music listener. When Bruce Springsteen and the Sessions Band appeared on the show as musical guests, O'Brien joined the 17-piece band, along with the Max Weinberg 7 and guests Jimmy Fallon and Thomas Haden Church, playing acoustic guitar and contributing backup vocals for the song "Pay Me My Money Down". On the June 13, 2008, episode of ''Late Night'', O'Brien simply walked onto the stage at the start of the show. Instead of his usual upbeat antics and monologue, O'Brien announced that he had just received news about the sudden death of his good friend, fellow NBC employee and frequent ''Late Night'' guest, Tim Russert. O'Brien proceeded to show two clips of his favorite Russert ''Late Night'' moments. On February 20, 2009, NBC aired the last episode of ''Late Night with Conan O'Brien''. The show consisted of a compilation of previous ''Late Night'' clips and included a surprise appearance by former sidekick, Andy Richter. Will Ferrell, John Mayer, and the White Stripes also appeared. O'Brien ended the episode by destroying the set with an axe, handing out the pieces of the set to the audience, and thanking a list of people who helped him get to that point in his career. Among those thanked were Lorne Michaels, David Letterman, Jay Leno, and O'Brien's wife and children.

The Colbert / O'Brien / Stewart mock feud

During the writers' strike in 2008, O'Brien staged a mock feud with Comedy Central's Jon Stewart (of ''The Daily Show'') and Stephen Colbert (of ''The Colbert Report'') over a dispute about which of the three were responsible for giving a "bump" to Mike Huckabee's campaign to become the Republican presidential nominee. This feud crossed over all three shows during the 2007–08 Writers Guild of America strike.

''The Tonight Show'' (2009–2010)

In 2004, O'Brien negotiated a new contract with NBC. As part of the deal, O'Brien would take over ''The Tonight Show'' from Jay Leno in 2009. O'Brien was a guest on Jay Leno's final episode of ''The Tonight Show''. On June 1, 2009, Will Ferrell became Conan's first ''Tonight Show'' guest on the couch and Pearl Jam appeared as his first musical guest.

During the taping of the Friday, September 25, 2009, episode of ''The Tonight Show'', O'Brien suffered from a mild concussion after he slipped and hit his head while running a race as part of a comedy sketch with guest Teri Hatcher. He was examined at a hospital and released the same day. A rerun was aired that night, but O'Brien returned to work the following Monday and poked fun at the incident.

Departure from ''The Tonight Show''

On Thursday, January 7, 2010, NBC executive Jeff Zucker met with Jay Leno and Conan O'Brien to discuss how to get Leno out of prime time, where his ratings were lackluster, and back into late night. A proposal was made that would see O'Brien remain as host of ''The Tonight Show'', which would be moved to 12:05 am with Leno hosting a 30-minute show at 11:35 pm On January 10, NBC Universal Television Entertainment Chairman Jeff Gaspin confirmed that ''The Jay Leno Show'' would indeed end at the start of the Winter Olympics on February 12, 2010, and be moved to 11:35 pm following the Olympics coverage. Sources familiar with the situation told the ''New York Post'' that O'Brien was unhappy with NBC's plan.

On January 12, O'Brien released this statement: "I sincerely believe that delaying ''The Tonight Show'' into the next day to accommodate another comedy program will seriously damage what I consider to be the greatest franchise in the history of broadcasting. ''The Tonight Show'' at 12:05 simply isn’t ''The Tonight Show.''" On January 21, 2010, it was announced that Conan had reached a deal with NBC that would see him exit ''The Tonight Show'' the next day. The deal also granted him $45 million, of which $12 million was designated for distribution to his staff, who had moved with Conan to Los Angeles from New York when he left ''Late Night''.

The final ''Tonight Show'' with Conan aired January 22, 2010, and featured guests Tom Hanks, Steve Carell (who did an exit interview and shredded Conan's ID badge), Neil Young (singing "Long May You Run"), and Will Ferrell. For Ferrell's appearance, Conan played guitar with the band and Ferrell sang "Free Bird" while reprising his ''SNL'' cowbell. Ferrell's wife, Viveca Paulin, together with Ben Harper, Beck, and ZZ Top guitarist Billy Gibbons, also joined the band for this final performance.

Jay Leno returned to ''The Tonight Show'' following NBC's coverage of the 2010 Winter Olympic Games. Under the $45 million deal with NBC, Conan was allowed to start working for another network as soon as September 2010. Conan's rumored next networks ranged anywhere from Fox to Comedy Central.

After ''Tonight''

On February 8, 2010, it was reported that O'Brien was attempting to sell his Central Park West penthouse in New York with an asking price of $35 million. He purchased the apartment in 2007 for $10 million. Two years earlier, O'Brien had purchased a home in the Brentwood section of Los Angeles for over $10.5 million. Some industry insiders have speculated that O'Brien had chosen to stay on the west coast in order to facilitate a return to late night television and because he did not want to put his children through another move.

On February 24, 2010, O'Brien attracted media attention for starting a Twitter account. His tweets, although primarily jokes, amounted to his first public statements since leaving ''The Tonight Show'' one month earlier. After about one hour, O'Brien's subscriber list had rocketed to over 30,000 members and approximately 30 minutes later, he was on the brink of passing 50,000 followers, already 20,000 more than the verified @jayleno account. After 24 hours, O'Brien had well over 300,000 followers. In late May 2010, he surpassed the one million mark for number of Twitter followers, and he has over 3.3 million followers.

O'Brien has been named to the 2010 Time 100, a list compiled by TIME of the 100 most influential people in the world as voted on by readers. After being prohibited from making television appearances of any kind until May, O'Brien spoke about the ''Tonight Show'' conflict on the CBS newsmagazine ''60 Minutes'' on May 2, 2010. During the interview with Steve Kroft, O'Brien said the situation felt "like a marriage breaking up suddenly, violently, quickly. And I was just trying to figure out what happened." He also said he "absolutely" expected NBC to give him more of a chance and that, if in Jay Leno's position, he would not have come back to ''The Tonight Show''. However, Conan said he did not feel he got shafted. "It's crucial to me that anyone seeing this, if they take anything away from this, it's I'm fine. I'm doing great," said O'Brien. "I hope people still find me comedically absurd and ridiculous. And I don't regret anything."

The Legally Prohibited from Being Funny on Television Tour

On March 11, 2010, O'Brien announced via his Twitter account that he would embark on a 30-city live tour beginning April 12, 2010, entitled "The Legally Prohibited from Being Funny on Television Tour". Co-host Andy Richter, along with members of the former Tonight Show Band, joined O'Brien on the tour. Max Weinberg, however, was not able to join, except for a guest appearance at one of Conan's New York City shows. VIP tickets for $695 offered the opportunity to meet O'Brien in person. On the same day, teamcoco.com—an apparently official website—was launched.

On April 12, 2010, O'Brien opened his two-month comedy tour in Eugene, Oregon, with a crowd of 2,500 and no TV cameras. The tour traveled through America's Northwest and Canada before moving on to larger cities, including Los Angeles and New York City, where he performed on the campuses that house both of the NBC-owned studios he formerly occupied. The tour ended in Atlanta on June 14. With ticket prices starting at $40, "The Legally Prohibited from Being Funny on Television Tour" was effectively sold out.

''Conan'' (2010–present)

On April 12, 2010, just hours before the start of his tour in Eugene, Oregon, O'Brien announced that he would host a new show on cable station TBS. The show debuted on November 8, 2010, and airs Monday through Thursday beginning at 11:00 pm ET/10:00 pm CT. O'Brien's addition moved ''Lopez Tonight'' with George Lopez back one hour to midnight ET/11:00 pm CT. Refusing at first to do to Lopez what had happened to him at NBC, O'Brien agreed to join the network after Lopez called to persuade him to come to TBS. In Canada, CTV will air the show and in Turkey, CNBC-e will air the show.

Other networks that were reportedly interested in O'Brien include TBS' sister networks TNT and HBO, Fox, FX, Comedy Central, Showtime, Revision3, and even the NBC Universal-owned USA Network.

On September 1, 2010, O'Brien announced via his Twitter account and Team Coco YouTube page that the title of his new show on TBS would simply be ''Conan''.

Television writer/producer (2002–present)

In 2004, O'Brien apologized to Canadians for engaging in Quebec bashing, something which some felt reflected prejudice against Francophones. On March 7, 2006, NBC announced a new adventure/comedy series entitled ''Andy Barker, P.I.'', starring O'Brien's former sidekick, Andy Richter. O'Brien was executive producer and co-wrote the pilot. After six episodes and low ratings, the show was canceled despite being named by ''Entertainment Weekly'' as one of the Top Ten Shows of 2007. USA Network has handed out a 90-minute, cast-contingent pilot order to the medical-themed ''Operating Instructions'' from O'Brien's production banner. O'Brien will serve as an executive producer through his Conaco label. The script comes from ''Just Shoot Me'' veterans Judd Pillot and John Peaslee who will also executive produce." NBC ordered two pilots from Conaco in January 2010, the one-hour courtroom drama, ''Outlaw'', and a half-hour comedy. ''Outlaw'' was later green-lit to series and premiered on September 15, 2010.

Voice work and guest appearances

O'Brien's first guest appearance after beginning his late-night career was on the show he used to write for, ''The Simpsons''. He played himself in the season five episode "Bart Gets Famous", interviewing Bart Simpson during his rise to fame as a catchphrase comedian. The episode was produced after O'Brien's audition to replace David Letterman, but before he was hired for the show. O'Brien recorded his part after his own show went on the air, though he believed his show would be canceled by the time "Bart Gets Famous" aired. In 2006, he voiced himself in a short ''South Park'' scene as part of the opening of the 2006 Emmy Awards. In 2005, he provided the voice of Robert Todd Lincoln in the audio book version of ''Assassination Vacation'' by Sarah Vowell.

O'Brien has made multiple voice appearances on the Adult Swim series ''Robot Chicken'', including the specials ''Robot Chicken: Star Wars'', and ''Robot Chicken: Star Wars Episode II'' as the voice of the bounty hunter Zuckuss. On the TV show ''30 Rock'', O'Brien is depicted as an ex-boyfriend of lead character Liz Lemon, who works in the same building. In the episode "Tracy Does Conan," Conan appears as himself, awkwardly reunited with Lemon and coerced by network executive Jack Donaghy into having the character Tracy Jordan on ''Late Night'', despite having been assaulted in Jordan's previous appearance.

O'Brien made an appearance on ''Futurama'' in the second-season episode "Xmas Story". O'Brien plays himself as a head in a jar and still alive in the year 3000. O'Brien performs a stand-up routine at a futuristic ski lodge while being heckled by Bender the robot.

O'Brien also made a cameo appearance on the U.S. version of ''The Office''. In the episode "Valentine's Day", Michael believes that he spots former ''SNL'' cast member, Tina Fey, but has actually mistaken another woman for her. In the meantime, Conan has a quick walk-on and the camera crew informs Michael, when he returns from talking to the Tina Fey lookalike.

In January 2010, O'Brien appeared in ''The Simpsons 20th Anniversary Special – In 3-D! On Ice!'' to honor the show he had written for in the early 1990s.

O'Brien created a superhero character with veteran DC Comics animator Bruce Timm during one episode of ''Conan''. Named "The Flaming C", the superhero bears a likeness to O'Brien, with a typically muscular superhero body and costume with chest insignia, but also with idiosyncrasies arbitrarily suggested by O'Brien like an oven mitt, a jai alai glove, marijuana leaf buckle, golf shoes, sock garters and fishnet stockings. O'Brien later aired a clip in which the character appears in ''Young Justice''.

Emmy host

O'Brien hosted the 58th Primetime Emmy Awards on August 27, 2006, to critical acclaim. He had previously hosted the Primetime Emmys in 2002, and co-hosted in 2003.

Commercials and product placements

According to Marketing Evolutions, the company that produces Q Scores measuring the familiarity and appeal of celebrities and brands, almost 74% of U.S. consumers are familiar with O'Brien. In spite of that high score, O'Brien hasn't done many commercials for corporate America; two recent examples are a 2009 appearance in a Super Bowl XLIII commercial for Bud Light, and one for American Express, which debuted on the same day as ''Conan''. He donated his proceeds from the Super Bowl commercial to the Fresh Air Fund, a charity which sends inner-city New York children to the country for vacations. According to ''The Wall Street Journal'', O'Brien was paid more than one million dollars to do the American Express commercial, which required him to spend three days in Jaipur, India filming it.

While O'Brien has done few commercials, he "does do plenty of promoting, weaving product pitches into his show"; he has said "it's increasingly incumbent to help with tie-ins [but] if it can't be funny, I'd rather go hungry."

Comedy and mannerisms

On ''Late Night,'' O'Brien became known for his active and spontaneous hosting style. He started off every show by saying, "We have a great show for you tonight" and ended nearly every show with a quick, quirky "Bye, everybody, bye!" His stage habits included, but were not limited to, mime, self-deprecation, dramatic expressions, various impressions, use of awkward pauses or responses, moving his hair and scalp back and forth and constantly buttoning and unbuttoning his suit jacket. He frequently made fun of and interacted with the audience and commonly made light of his own appearance, including his hairstyle, his pale skin, his clumsiness, and his height. He continued many of these themes into his position as host of ''The Tonight Show''.

One of O'Brien's trademarks is to perform the "string dance." He also does intentionally poor and exaggerated impressions of celebrities that are often reduced to a specific characteristic, phrase, or gesture that represents that person.

Personal life

O'Brien met Elizabeth Ann 'Liza' Powel in 2000 when she appeared on ''Late Night with Conan O'Brien'' in an advertising skit involving Foote, Cone & Belding, where she worked as senior copywriter. The couple dated for nearly 18 months before their 2002 marriage in Powel's hometown of Seattle. O'Brien and Powel have a daughter, Neve (born in 2003) and a son, Beckett (born in 2005).

O'Brien repeatedly affirms his Irish Catholic heritage on his show. On a 2009 episode of ''Inside the Actors Studio'', he stated that both sides of his family moved to America from Ireland in the 1850s, subsequently marrying only other Irish Catholics, and that his lineage is thus 100% Irish Catholic.

He has been a staunch Democrat since casting his first vote for President in 1984 for Walter Mondale, although he considers himself a moderate on the political spectrum. O'Brien's longtime friend and former roommate at Harvard is Father Paul B. O' Brien, with whom he founded ''Labels Are For Jars'', an antihunger organization based in Lawrence, Massachusetts, and helped open the ''Cor Unum'' meal center in 2006. The two are not related.

In January 2008, after his show was put on hold for two months owing to the strike by the Writers Guild of America, he reemerged on late-night TV sporting a beard, which guest Tom Brokaw described as making him look like "a draft dodger from the Civil War." After leaving ''The Tonight Show with Conan O'Brien'' on Jan. 22, 2010, O'Brien again grew a beard, which he kept until May 2, 2011, when it was partially shaved on the set of his TBS talk show, ''Conan'', by Will Ferrell with battery-operated clippers (and completely shaved off-screen by a professional barber). The event was dubbed on the show as "Beardocalypse," and included a contest for fan-submitted artwork.

O'Brien purchased a $10.5-million mansion in Brentwood, Los Angeles, California, to prepare for his move there in 2009 from New York City to host ''The Tonight Show'' at Universal Studios Hollywood. As part of a long running gag, he brought his 1992 Ford Taurus SHO with him to California, showcasing it on both the inaugural episodes of ''The Tonight Show'' and ''Conan''.

Victim of stalking

It was reported that since September 2006, O'Brien had allegedly been stalked by Father David Ajemian of the Archdiocese of Boston, who, despite multiple warnings to stop, had been sending O'Brien letters signed as "your priest stalker" and coming in contact with O'Brien's parents. Frustrated that he had been denied a spot in the ''Late Night'' audience, Ajemian sent a letter to O'Brien stating that he flew to New York "in the dimming hope that you might finally acknowledge me." He stated in another letter, "Is this the way you treat your most dangerous fans??? You owe me big time, pal." In another letter, Ajemian seemed to make a death threat, saying, "Remember Frank Costello once dodged a bullet in your building and so can you." Ajemian then tried to forcefully enter a taping of ''Late Night'' but was caught and arrested. He was previously warned by the NBC security team to stay away from the studio. After a psychological evaluation, he was deemed fit to stand trial. He has since been bailed out of jail. He was then reported missing by his father on November 10, 2007. He was found and underwent evaluation at a hospital. It is known that the two had attended Harvard University at the same time. He was found fit to stand trial on April 4, 2008. On April 8, 2008, Ajemian pleaded guilty to stalking, stating that he "never meant to cause anxiety or to upset anyone." He was ordered to pay a $95 US$ court charge and to sign a two-year restraining order barring him from coming near O'Brien. On September 11, 2008, Ajemian checked himself out of his treatment at a hospital against the wishes of his bishop, Seán Patrick O'Malley. Cardinal O'Malley then released a statement saying that because he violated his bishop's wishes, Ajemian could no longer serve as a priest in the Catholic Church. As of late 2010, Ajemian has moved on to stalking Anthony Everett.

Nicknames

Conan acquired the nickname "Coco" after its use in the first "Twitter Tracker" sketch during the second episode of his ''Tonight Show'' run. Guest Tom Hanks used the nickname during his subsequent interview, even getting the audience to chant it. In reaction to the moniker, Conan remarked to Hanks in jest, "If that catches on, I'll sue you."

In a March 23, 2011, interview with WWE Champion The Miz on ''Conan'', The Miz dubbed Conan "The Ginja Ninja", a reference to Conan's red hair and the fact that he came back fighting to get his new late-night talk show. A week later, "Team Ginja Ninja" T-shirts were available on TeamCoco.com.

Filmography

;Appearances on talk shows: Guest appearance on ''The Tonight Show with Jay Leno'' (1993, 2007, 2008, 2009)

Guest appearance on ''Late Night with David Letterman'' (1993)

Guest appearance on ''The Jon Stewart Show'' (1993)

''Late Night with Conan O'Brien'' (1993–2009)

Guest appearance on ''Last Call with Carson Daly'' (2002)

Guest appearance on ''The Daily Show with Jon Stewart'' (2007, 2008)

Guest appearance on ''The Colbert Report'' (2007, 2008)

''Late Night with Jimmy Fallon'' (2009)

''The Tonight Show with Conan O'Brien'' (2009–2010)

Guest appearance on ''Lopez Tonight'' (2010, 2011)

''Conan'' (November 8, 2010–)

;Other shows:

''Saturday Night Live'' (1991, 2001)

''The Simpsons'' (''as himself'') (1994)

''Space Ghost: Coast to Coast'' (''as himself'') (1999)

''Mr. Show with Bob and David'' (1995)

''The Single Guy'' (1996)

''Arli$$'' (1996)

''Spin City'' (1999)

''Futurama'' (1999)

''Storytelling'' (2001)

''Vanilla Sky'' (2001)

''Sugar & Spice'' (2001) ''(cameo in end credits)''

''Dr. Katz'' (''as himself'') (2002)

''Andy Richter Controls the Universe'' (2003)

''End of the Century'' (2003)

''Bewitched'' (''as himself'') (2005)

''Robot Chicken'' (2005)

''The Denial Twist (The White Stripes music video)'' (2005)

''O'Grady'' (2006)

''The Office'' ''(cameo)'' (2006)

''Queer Duck'' (''as himself'') (2006)

''30 Rock'' (''as himself'') (2007, 2009)

Robot Chicken Star Wars (as ''Zuckuss'') (2007)

''Sesame Street'' (2008)

''Robot Chicken: Star Wars Episode II'' (as ''Zuckuss'', Star Wars nerd) (2008)

''The Backyardigans'' (2009)

''60 Minutes'' (interview) (2010)

''Eagleheart'' (himself) (2010)

Awards and nominations

Year	Award	Work	Category	Result
1989	Emmy Award		Outstanding Writing in a Variety or Music Program
1990	Emmy Award	Outstanding Writing in a Variety or Music Program
1991	Emmy Award	Outstanding Writing in a Variety or Music Program
1996	Emmy Award		Outstanding Individual Achievement in Writing for a Variety or Music Program
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
1998	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
People's Choice Award	Favorite Late Night Talk Show Host
Telvis Award	For the color spot of the year	Special Telvis
Writers Guild of America Award		Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
People's Choice Award	Favorite Late Night Talk Show Host
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
2008	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
	Emmy Award	Outstanding Individual Achievement in Writing for a Variety or Music Program
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
			Outstanding Comedy, Music or Variety Series
Outstanding Writing for a Variety, Music or Comedy Series
Writers Guild of America Award	Comedy/Variety (Including Talk) – Series
	People's Choice Award	rowspan="4"	Favorite TV Talk Show Host
	Outstanding Comedy, Music or Variety Series
Outstanding Writing for a Variety, Music or Comedy Series
Outstanding Lighting Design/Lighting Direction for a Variety, Music or Comedy Series
American Express	Outstanding Commercial

References

Further reading

External links

in 2006 in 1998

TMZ page on Conan O'Brien

I'm With Coco Facebook

Category:1963 births Category:Living people Category:American comedians Category:American television talk show hosts Category:American television writers Category:Emmy Award winners Category:The Groundlings Category:Harvard Lampoon people Category:Harvard University alumni Category:American comedians of Irish descent Category:American people of Irish descent Category:American writers of Irish descent Category:Late night television talk show hosts Category:Late Night with Conan O'Brien Category:People from Brookline, Massachusetts Category:Writers from Massachusetts Category:Massachusetts Democrats

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name	Sir David Attenborough
birth date	May 08, 1926
birth place	Isleworth, London, England
residence	Richmond, London
nationality	British
alma mater
occupation
title
spouse	Jane Elizabeth Ebsworth Oriel (m. 1950–1997, her death)
children
footnotes	}}

Sir David Frederick Attenborough () OM, CH, CVO, CBE, FRS, FZS, FSA (born 8 May 1926 in London, England) is a broadcaster and naturalist. His career as the respected face and voice of natural history programmes has endured for more than 50 years. He is best known for writing and presenting the nine ''Life'' series, in conjunction with the BBC Natural History Unit, which collectively form a comprehensive survey of all life on the planet. He is also a former senior manager at the BBC, having served as controller of BBC Two and director of programming for BBC Television in the 1960s and 1970s.

He is a younger brother of director, producer and actor Richard Attenborough.

Early life and family

Attenborough grew up in College House on the campus of the University of Leicester, where his father, Frederick, was principal. He is the middle of three sons (his elder brother, Richard, became an actor/film director and his younger brother, John Michael Attenborough, an executive at Alfa Romeo). During World War II his parents also adopted two Jewish refugee girls from Europe.

Attenborough spent his childhood collecting fossils, stones and other natural specimens. He received encouragement in this pursuit at age seven, when a young Jacquetta Hawkes admired his "museum". A few years later, one of his adoptive sisters gave him a piece of amber filled with prehistoric creatures; some 50 years later, it would be the focus of his programme ''The Amber Time Machine''.

Attenborough was educated at Wyggeston Grammar School for Boys in Leicester and then won a scholarship to Clare College, Cambridge in 1945, where he studied geology and zoology and obtained a degree in Natural Sciences. In 1947, he was called up for National Service in the Royal Navy and spent two years stationed in North Wales and the Firth of Forth.

In 1950, Attenborough married Jane Elizabeth Ebsworth Oriel; the marriage lasted until her death in 1997. The couple had two children, Robert and Susan.

His son, Robert Attenborough, is a senior lecturer in Bioanthropology for the School of Archaeology and Anthropology at the Australian National University in Canberra.

First years at BBC

After leaving the Navy, Attenborough took a position editing children's science textbooks for a publishing company. He soon became disillusioned with the work, however, and in 1950, he applied for a job as a radio talks producer with the BBC. Although he was rejected for this job, his CV later attracted the interest of Mary Adams, head of the Talks (factual broadcasting) department of the BBC's fledgling television service. Attenborough, like most Britons at that time, did not own a television, and he had seen only one programme in his life. However, he accepted Adams' offer of a three-month training course, and in 1952, he joined the BBC full time. Initially discouraged from appearing on camera because Adams thought his teeth were too big, he became a producer for the Talks Department, which handled all non-fiction broadcasts. His early projects included the quiz show ''Animal, Vegetable, Mineral?'' and ''Song Hunter'', a series about folk music presented by Alan Lomax.

Attenborough's association with natural history programmes began when he produced and presented the three-part series ''The Pattern of Animals''. The studio-bound programme featured animals from London Zoo, with the naturalist Sir Julian Huxley discussing their use of camouflage, aposematism and courtship displays. Through this programme, Attenborough met Jack Lester, the curator of the zoo's reptile house, and they decided to make a series about an animal-collecting expedition. The result was ''Zoo Quest'', first broadcast in 1954, which Attenborough presented at short notice, due to Lester being taken ill.

In 1957, the BBC Natural History Unit was formally established in Bristol. Attenborough was asked to join it, but declined, not wishing to move from London where he and his young family were settled. Instead he formed his own department, the Travel and Exploration Unit, which allowed him to continue to front the ''Zoo Quest'' programmes as well as produce other documentaries, notably the ''Travellers’ Tales'' and ''Adventure'' series.

In the early 1960s, Attenborough resigned from the permanent staff of the BBC to study for a postgraduate degree in social anthropology at the London School of Economics, interweaving his study with further filming. However, he accepted an invitation to return to the BBC as Controller of BBC Two before he could finish the degree.

BBC administration

Attenborough became the Controller of BBC Two in March 1965, but had a clause inserted in his contract that would allow him to continue making programmes on an occasional basis. Later the same year, he filmed elephants in Tanzania, and in 1969, he made a three-part series on the cultural history of Bali. For the 1971 film ''A Blank on the Map'', he joined the first Western expedition to a remote highland valley in New Guinea to seek out an uncontacted tribe.

BBC Two was launched in 1964, but had struggled to capture the public's imagination. When Attenborough arrived as Controller, he quickly abolished the channel's quirky kangaroo mascot and shook up the schedule. With a mission to make BBC Two's output diverse and different from that offered by other networks, he began to establish a portfolio of programmes that defined the channel's identity for decades to come. Under his tenure, music, the arts, entertainment, archaeology, experimental comedy, travel, drama, sport, business, science and natural history all found a place in the weekly schedules. Often, an eclectic mix was offered within a single evening's viewing. Programmes he commissioned included ''Man Alive'', ''Call My Bluff'', ''Chronicle'', ''Life'', ''One Pair of Eyes'', ''The Old Grey Whistle Test'', ''Monty Python's Flying Circus'' and ''The Money Programme''. When BBC Two became the first British channel to broadcast in colour in 1967, Attenborough took advantage by introducing televised snooker, as well as bringing rugby league to British television on a regular basis via the BBC2 Floodlit Trophy.

One of his most significant decisions was to order a 13-part series on the history of Western art, to show off the quality of the new UHF colour television service that BBC Two offered. Broadcast to universal acclaim in 1969, ''Civilisation'' set the blueprint for landmark authored documentaries, which were informally known as "tombstone" or "sledgehammer" projects. Others followed, including Jacob Bronowski's ''The Ascent of Man'' (also commissioned by Attenborough), and Alistair Cooke's ''America''. Attenborough thought that the story of evolution would be a natural subject for such a series. He shared his idea with Chris Parsons, a producer at the Natural History Unit, who came up with the title ''Life on Earth'' and returned to Bristol to start planning the series. Attenborough harboured a strong desire to present the series himself, but this would not be possible so long as he remained in a management post.

In 1969, Attenborough was promoted to Director of Programmes, making him responsible for the output of both BBC channels. His tasks, which included agreeing budgets, attending board meetings and firing staff, were now far removed from the business of filming programmes. When Attenborough's name was being suggested as a candidate for the position of Director General of the BBC in 1972, he phoned his brother Richard to confess that he had no appetite for the job. Early the following year, he left his post to return to full-time programme making, leaving him free to write and present the planned natural history epic.

Return to broadcasting

After his resignation, Attenborough became a freelance broadcaster and immediately started work on his next project, a pre-arranged trip to Indonesia with a crew from the Natural History Unit. It resulted in the 1973 series ''Eastwards with Attenborough'', which was similar in tone to the earlier ''Zoo Quests'' but without the animal-collecting element.

On his return, he began to work on the scripts for ''Life on Earth''. Due to the scale of his ambition, the BBC decided to partner with an American network to secure the necessary funding. While the negotiations were proceeding he worked on a number of other television projects. He presented a series on tribal art (''The Tribal Eye'', 1975) and another on the voyages of discovery (''The Explorers'', 1975). He also presented a BBC children's series about cryptozoology entitled ''Fabulous Animals'' (1975), which featured mythical creatures such as the griffin and kraken. Eventually, the BBC signed a co-production deal with Turner Broadcasting and ''Life on Earth'' moved into production in 1976.

"Life" series

Beginning with ''Life on Earth'' in 1979, Attenborough set about creating a body of work which became a benchmark of quality in wildlife film-making and influenced a generation of documentary film-makers. The series also established many of the hallmarks of the BBC's natural history output. By treating his subject seriously and researching the latest discoveries, Attenborough and his production team gained the trust of the scientific community, who responded by allowing him to feature their subjects in his programmes. In Rwanda, for example, Attenborough and his crew were granted privileged access to film Dian Fossey's research group of mountain gorillas. Innovation was another factor in ''Life on Earth'''s success: new film-making techniques were devised to get the shots Attenborough wanted, with a focus on events and animals that were hitherto unfilmed. Computerised airline schedules, which had only recently been introduced, enabled the series to be elaborately devised so that Attenborough visited several locations around the globe in each episode, sometimes even changing continents mid-sentence. Although appearing as the on-screen presenter, he consciously restricted his pieces to camera to give his subjects top billing.

The success of ''Life on Earth'' prompted the BBC to consider a follow-up, and five years later, ''The Living Planet'' was screened. This time, Attenborough built his series around the theme of ecology, the adaptations of livings things to their environment. It was another critical and commercial success, generating huge international sales for the BBC. In 1990, ''The Trials of Life'' completed the original "Life" trilogy, looking at animal behaviour through the different stages of life. The series drew strong reactions from the viewing public for its sequences of killer whales hunting sea lions on a Patagonian beach and chimpanzees hunting and violently killing a colobus monkey.

In the 1990s, Attenborough continued to use the "Life" moniker for a succession of authored documentaries. In 1993, he presented ''Life in the Freezer'', the first television series to survey the natural history of Antarctica. Although past normal retirement age, he then embarked on a number of more specialised surveys of the natural world, beginning with plants. They proved a difficult subject for his producers, who had to deliver five hours of television featuring what are essentially immobile objects. The result, ''The Private Life of Plants'' (1995), showed plants as dynamic organisms by using time-lapse photography to speed up their growth.

Prompted by an enthusiastic ornithologist at the BBC Natural History Unit, Attenborough then turned his attention to the animal kingdom and in particular, birds. As he was neither an obsessive twitcher, nor a bird expert, he decided he was better qualified to make ''The Life of Birds'' (1998) on the theme of behaviour. The order of the remaining "Life" series was dictated by developments in camera technology. For ''The Life of Mammals'' (2002), low-light and infrared cameras were deployed to reveal the behaviour of nocturnal mammals. The series contains a number of memorable two shots of Attenborough and his subjects, which included chimpanzees, a blue whale and a grizzly bear. Advances in macro photography made it possible to capture natural behaviour of very small creatures for the first time, and in 2005, ''Life in the Undergrowth'' introduced audiences to the world of invertebrates.

At this point, Attenborough realised that he had spent 20 years unconsciously assembling a collection of programmes on all the major groups of terrestrial animals and plants — only reptiles and amphibians were missing. When ''Life in Cold Blood'' was broadcast in 2008, he had the satisfaction of completing the set, brought together in a DVD encyclopaedia called ''Life on Land''. In an interview that year, Attenborough was asked to sum up his achievement, and responded:

However, in 2010 Attenborough asserted that his forthcoming ''First Life'' — dealing with evolutionary history before ''Life on Earth'' — should also be included within the "Life" series. In the documentary ''Attenborough's Journey'' he stated, "This series, to a degree which I really didn't fully appreciate until I started working on it, really completes the set."

Other documentaries

Alongside the "Life" series, Attenborough has continued to work on other television documentaries, mainly in the natural history genre. He wrote and presented a series on man's influence on the natural history of the Mediterranean basin, ''The First Eden'', in 1987. Two years later, he demonstrated his passion for fossils in ''Lost Worlds Vanished Lives''.

Attenborough narrated every episode of ''Wildlife on One'', a BBC One wildlife series which ran for 253 episodes between 1977 and 2005. At its peak, it drew a weekly audience of eight to ten million, and the 1987 episode "Meerkats United" was voted the best wildlife documentary of all time by BBC viewers. He has also narrated over 50 episodes of ''Natural World'', BBC Two's flagship wildlife series. (Its forerunner, ''The World About Us'', was created by Attenborough in 1969, as a vehicle for colour television.) In 1997, he narrated the ''BBC Wildlife Specials'', each focussing on a charismatic species, and screened to mark the Natural History Unit's 40th anniversary.

As a writer and narrator, he continued to collaborate with the BBC Natural History Unit in the new millennium. Alastair Fothergill, a senior producer with whom Attenborough had worked on ''The Trials of Life'' and ''Life in the Freezer'', was making ''The Blue Planet'' (2001), the Unit's first comprehensive series on marine life. He decided not to use an on-screen presenter due to difficulties in speaking to camera through diving apparatus, but asked Attenborough to narrate the films. The same team reunited for ''Planet Earth'' (2006), the biggest nature documentary ever made for television, and the first BBC wildlife series to be shot in high definition. In 2009, Attenborough wrote and narrated ''Life'', a ten-part series focussing on extraordinary animal behaviour, and narrated ''Nature's Great Events'', which showed how seasonal changes trigger major natural spectacles.

By the turn of the millennium, Attenborough's authored documentaries were adopting a more overtly environmentalist stance. In ''State of the Planet'' (2000), he used the latest scientific evidence and interviews with leading scientists and conservationists to assess the impact of man's activities on the natural world. He later turned to the issues of global warming (''The Truth about Climate Change'', 2006) and human population growth (''How Many People Can Live on Planet Earth?'', 2009). He also contributed a programme which highlighted the plight of endangered species to the BBC's ''Saving Planet Earth'' project in 2007, the 50th anniversary of the Natural History Unit.

Interestingly, although Attenborough's documentaries have attained immense popularity in the United States, several have never been made available on DVD in NTSC format, most notably those that cast doubt upon conservative religious or political positions. These include:

''Life on Earth'', which examines the evidence for evolution.

''State of the Planet''

''The Truth About Climate Change''

''First Life'', which examines the origin of life.

Current projects

Attenborough continues to work into his ninth decade, and is currently involved in a number of projects for the BBC and for Sky.

He is writing and presenting ''Frozen Planet'', a major new series for BBC One which examines the impact of a warming climate on the people and wildlife of the polar regions. He has also recently completed two projects for BBC Two. ''Madagascar'' (which first aired weekly between the 9 to 23 February 2011) a three-part series giving an overview of Madagascar's unique wildlife. The accompanying documentary ''Attenborough and the Giant Egg'' (which first aired on the 2nd of March 2011) features the elephant bird egg which Attenborough discovered on his first filming expedition to the island in the 1960s.

Attenborough is also forging a new partnership with Sky, working on documentaries for the broadcaster's new 3D network, Sky 3D. Their first collaboration is ''Flying Monsters 3D'', a film about pterosaurs which debuted on Christmas Day 2010. A second film, ''Penguin Island 3D'', has also been announced. Both are produced by Atlantic Productions, the company behind Attenborough's 2010 series ''First Life''.

Other work

From 1983, Attenborough worked on two environmentally themed musicals with the WWF and writers Peter Rose and Anne Conlon. ''Yanomamo'' was the first, about the Amazon rainforest, and the second, ''Ocean World'', premiered at the Royal Festival Hall in 1991. They were both narrated by Attenborough on their national tour, and recorded on to audio cassette. ''Ocean World'' was also filmed for Channel 4 and later released.

In 1990, he highlighted the case of Mahjoub Sharif as part of the BBC's ''Prisoners of Conscience'' series.

In January 2009, the BBC commissioned Attenborough to provide a series of 20 ten-minute monologues covering the history of nature. Entitled ''David Attenborough's Life Stories'', they are broadcast on Radio 4 in the Friday night slot vacated by Alistair Cooke's ''Letter from America''. Part of Radio 4's ''A Point of View'' strand, the talks are also available as podcasts.

He appeared in the 2009 Children's Prom at the BBC Promenade Concerts and in the Last Night of the Proms on 12 September 2009, playing a floor polisher in Sir Malcolm Arnold's "A Grand, Grand Overture" (after which he was "shot" by Rory Bremner, who was playing the gun).

Attenborough also serves on the advisory board of ''BBC Wildlife'' magazine.

Achievements, awards and recognition

{| style="float:right; background:#f9f9f9; border:1px solid darkgrey;" |- !align="left"|Styles and honours |- |

Mr David Attenborough (1926–1974)

Mr David Attenborough CBE (1974–1983)

Mr David Attenborough CBE FRS (1983–1985)

Sir David Attenborough CBE FRS (1985–1991)

Sir David Attenborough CVO CBE FRS (1991–1996)

Sir David Attenborough CH CVO CBE FRS (1996–2005)

Sir David Attenborough OM CH CVO CBE FRS (2005–)

|}

Attenborough's contribution to broadcasting and wildlife film-making has brought him international recognition. He has been called "the great communicator, the peerless educator" and "the greatest broadcaster of our time". His programmes are often cited as an example of what public service broadcasting should be, even by critics of the BBC, and has influenced a generation of wildlife film-makers.

Honorary titles

In 2010 it was reported that Attenborough has been given 29 honorary degrees by British universities, more than any other person. In 1980, he was honoured by the Open University, with whom he has had a close association throughout his career. He also has honorary Doctor of Science awards from the University of Cambridge (1984) and the University of Oxford (1988). In 2006, the two eldest Attenborough brothers returned to their home city to receive the title of Distinguished Honorary Fellows of the University of Leicester, "in recognition of a record of continuing distinguished service to the University." David Attenborough was previously awarded an honorary Doctor of Letters degree by the university in 1970, and was made an honorary Freeman of the City of Leicester in 1990.

Attenborough has received the title Honorary Fellow from Clare College, Cambridge (1980), the Zoological Society of London (1998), the Linnean Society (1999), the Institute of Biology (2000) and the Society of Antiquaries (2007).

Recognition

Attenborough has featured as the subject of a number of BBC television programmes. ''Life on Air'' (2002) examined the legacy of his work and ''Attenborough the Controller'' (2002) focused on his time in charge of BBC Two. He also featured prominently in ''The Way We Went Wild'' (2004), a series about natural history television presenters, and ''100 Years of Wildlife Films'' (2007), a special programme marking the centenary of the nature documentary. In 2006, British television viewers were asked to vote for their ''Favourite Attenborough Moments'' for a UKTV poll to coincide with the broadcaster's 80th birthday. The winning clip showed Attenborough observing the mimicry skills of the superb lyrebird.

Attenborough was named as the most trusted celebrity in Britain in a 2006 Reader's Digest poll,. and the following year he won ''The Culture Show'''s Living Icon Award. He has also been named among the 100 Greatest Britons in a 2002 BBC poll and is one of the top ten "Heroes of Our Time" according to ''New Statesman'' magazine.

He has the distinction of having a number of newly-discovered species and fossils being named in his honour. In 1993, after discovering that the Mesozoic reptile ''Plesiosaurus conybeari'' had not, in fact, been a true plesiosaur, the paleontologist Robert Bakker renamed the species ''Attenborosaurus conybeari''. A fossilised armoured fish discovered at the Gogo Formation, Western Australia in 2008 was given the name ''Materpiscis attenboroughi'', after Attenborough had filmed at the site and highlighted its scientific importance in ''Life on Earth''. The ''Materpiscis'' fossil is believed to be the earliest organism capable of internal fertilisation.

He has also lent his name to a species of Ecuadorian flowering tree, ''Blakea attenboroughi'', one of the world's largest carnivorous plants, ''Nepenthes attenboroughii'', and one of only four species of long-beaked echidna, the critically endangered ''Zaglossus attenboroughi'', discovered by explorer and zoologist Tim Flannery in the Cyclops Mountains of New Guinea in 1998.

In September 2009, London's Natural History Museum opened the Attenborough Studio, part of its Darwin Centre development.

Awards

1970: BAFTA Desmond Davis Award

1974: Commander of the Order of the British Empire (CBE)

1980: BAFTA Fellowship

1983: Fellow of the Royal Society (FRS)

1985: Knighthood

1991: Commander of the Royal Victorian Order (CVO) for producing Queen Elizabeth II's Christmas broadcast for a number of years from 1986

1991: Foreign Honorary Member of the American Academy of Arts and Sciences

1996: Companion of Honour (CH) "for services to nature broadcasting"

1998: International Cosmos Prize

2003: Michael Faraday Prize awarded by the Royal Society

2004: Descartes Prize for Outstanding Science Communication Actions

2004: Caird Medal of the National Maritime Museum

2005: Order of Merit (OM)

2005: Nierenberg Prize for Science in the Public Interest

2006: National Television Awards Special Recognition Award

2006: Institute of Ecology and Environmental Management - Institute Medal in recognition of his outstanding contribution to the public perception and understanding of ecology

2006: The Culture Show British Icon Award

2007: British Naturalists' Association Peter Scott Memorial Award

2009: BAFTA Specialist factual award for ''Life In Cold Blood''

2009: Prince of Asturias Award

2010: Fonseca Prize

2010: Queensland Museum Medal

Views and advocacy

Environmental causes

Attenborough's programmes have often included references to the impact of human society on the natural world. The last episode of ''The Living Planet'', for example, focuses almost entirely on humans' destruction of the environment and ways that it could be stopped or reversed. Despite this, he has been criticised for not giving enough prominence to environmental messages. Some environmentalists feel that programmes like Attenborough's give a false picture of idyllic wilderness and do not do enough to acknowledge that such areas are increasingly encroached upon by humans.

However, his closing message from ''State of the Planet'' was forthright:

The future of life on earth depends on our ability to take action. Many individuals are doing what they can, but real success can only come if there's a change in our societies and our economics and in our politics. I've been lucky in my lifetime to see some of the greatest spectacles that the natural world has to offer. Surely we have a responsibility to leave for future generations a planet that is healthy, inhabitable by all species.

Attenborough has subsequently become more vocal in his support of environmental causes. In 2005 and 2006, he backed a BirdLife International project to stop the killing of albatross by longline fishing boats. He gave public support to WWF's campaign to have 220,000 square kilometres of Borneo's rainforest designated a protected area. He also serves as a vice-president of BTCV, vice-president of Fauna and Flora International, president of Butterfly Conservation and president of Leicestershire and Rutland Wildlife Trust. In 2003 he launched an appeal to create a rainforest reserve in Ecuador in memory of Christopher Parsons, the producer of ''Life on Earth'' and a personal friend, who had died the previous year. The same year, he helped to launch ARKive, a global project instigated by Parsons to gather together natural history media into a digital library. ARKive is an initiative of Wildscreen, of which Attenborough is a patron. He later became patron of the World Land Trust, and an active supporter. He supported Glyndebourne in their successful application to obtain planning permission for a wind turbine in an Area of Outstanding Natural Beauty, and gave evidence at the planning inquiry arguing in favour of the proposal.

Attenborough has repeatedly said that he considers human overpopulation to be the root cause of many environmental problems. In ''The Life of Mammals'', he made a plea for humans to curb population growth so that other species will not be crowded out. In 2009, he became a patron of Population Matters, (formerly known as the Optimum Population Trust), a UK charity advocating sustainable human populations.

He has written and spoken publicly about the fact that, despite past scepticism, he believes the Earth's climate is warming in a way that is cause for concern, and that this can likely be attributed to human activity. He summed up his thoughts at the end of his 2006 documentary "Can We Save Planet Earth?" as follows:

In the past, we didn't understand the effect of our actions. Unknowingly, we sowed the wind and now, literally, we are reaping the whirlwind. But we no longer have that excuse: now we do recognise the consequences of our behaviour. Now surely, we must act to reform it — individually and collectively, nationally and internationally — or we doom future generations to catastrophe.

In a 2005 interview with ''BBC Wildlife'' magazine, Attenborough said he considered George W. Bush to be the era's top "environmental villain". In 2007, he further elaborated on the USA's consumption of energy in relation to its population. When asked if he thought America to be "the villain of the piece", he responded:

I don't think whole populations are villainous, but Americans are just extraordinarily unaware of all kinds of things. If you live in the middle of that vast continent, with apparently everything your heart could wish for just because you were born there, then why worry? [...] If people lose knowledge, sympathy and understanding of the natural world, they're going to mistreat it and will not ask their politicians to care for it.

Religion and creationism

In a December 2005 interview with Simon Mayo on BBC Radio Five Live, Attenborough stated that he considers himself an agnostic. When asked whether his observation of the natural world has given him faith in a creator, he generally responds with some version of this story, making reference to the ''Loa loa'' parasitic worm:

My response is that when Creationists talk about God creating every individual species as a separate act, they always instance hummingbirds, or orchids, sunflowers and beautiful things. But I tend to think instead of a parasitic worm that is boring through the eye of a boy sitting on the bank of a river in West Africa, [a worm] that's going to make him blind. And [I ask them], 'Are you telling me that the God you believe in, who you also say is an all-merciful God, who cares for each one of us individually, are you saying that God created this worm that can live in no other way than in an innocent child's eyeball? Because that doesn't seem to me to coincide with a God who's full of mercy'.

He has explained that he feels the evidence all over the planet clearly shows evolution to be the best way to explain the diversity of life, and that "as far as I'm concerned, if there is a supreme being then he chose organic evolution as a way of bringing into existence the natural world."

In a BBC Four interview with Mark Lawson, Attenborough was asked if he at any time had any religious faith. He replied simply, "No." He has also said "It never really occurred to me to believe in God".

In 2002, Attenborough joined an effort by leading clerics and scientists to oppose the inclusion of creationism in the curriculum of UK state-funded independent schools which receive private sponsorship, such as the Emmanuel Schools Foundation. In 2009, Attenborough stated that the Book of Genesis, by saying that the world was there for people to dominate, had taught generations that they can "dominate" the environment, and that this has resulted in the devastation of vast areas of the environment. Attenborough further explained to the science journal ''Nature'', "That's why Darwinism, and the fact of evolution, is of great importance, because it is that attitude which has led to the devastation of so much, and we are in the situation that we are in."

Also in early 2009, the BBC broadcast an Attenborough one-hour special, ''Charles Darwin and the Tree of Life''. In reference to the programme, Attenborough stated that "People write to me that evolution is only a theory. Well, it is not a theory. Evolution is as solid a historical fact as you could conceive. Evidence from every quarter. What is a theory is whether natural selection is the mechanism and the only mechanism. That is a theory. But the historical reality that dinosaurs led to birds and mammals produced whales, that's not theory." He strongly opposes creationism and its offshoot "intelligent design", saying that a survey that found a quarter of science teachers in state schools believe that creationism should be taught alongside evolution in science lessons was "really terrible".

In March 2009, Attenborough appeared on ''Friday Night with Jonathan Ross''. Attenborough stated that he felt evolution did not rule out the existence of a God and accepted the title of agnostic saying, "My view is: I don't know one way or the other but I don't think that evolution is against a belief in God."

BBC and public service broadcasting

Attenborough is a lifelong supporter of the BBC, public broadcasting and the television licence. He has said:

PSB, to me, is not about selecting individual programme strands here or there, financing them from some outside source and then foisting them upon commercial networks. Public Service Broadcasting, watched by a healthy number of viewers, with programmes financed in proportion to their intrinsic needs and not the size of the audience, can only effectively operate as a network — a network whose aim is to cater for the broadest possible range of interests, popular as well as less popular, a network that measures its success not only by its audience size but by the range of its schedule.

Public service broadcasting is one of the things that distinguishes this country and makes me want to live here. I have spent all my life in it. I would be very distressed if public service broadcasting was weakened. I have been at the BBC since 1952, and know the BBC is constantly being battered. It is today.

If you could demonstrate that the BBC was grossly extravagant there might be a case for saying OK take it away. But in fact the BBC per minute in almost every category is as cheap as you can find anywhere in the world and produces the best quality. If you take the money away, which part of the BBC will you remove? The BBC has gone through swingeing staff cuts. It has been cut to the bone, if you divert licence fee money elsewhere, you cut quality and services. There is always that threat from politicians who will say your licence fee is up for grabs. We will take it. There is a lot of people who want to see the BBC weakened. They talk of this terrible tax of the licence fee. Yet it is the best bargain that is going. Four radio channels and god knows how many TV channels. It is piffling.

There have always been politicians or business people who have wanted to cut the BBC back or stop it saying the sort of things it says. There's always been trouble about the licence and if you dropped your guard you could bet our bottom dollar there'd be plenty of people who'd want to take it away. The licence fee is the basis on which the BBC is based and if you destroy it, broadcasting... becomes a wasteland.

Attenborough expressed regret at some of the changes made to the BBC in the 1990s by Director-General John Birt, who introduced an internal market at the corporation, slimmed and even closed some departments and outsourced much of the corporation’s output to private production companies, in line with the Broadcasting Act 1990. He has said:

There is no question but that Birtism . . . has had some terrible results. On the other hand, the BBC had to change. Now it has to produce programmes no one else can do. Otherwise, forget the licence fee.

The Bristol Unit has suffered along with the rest of the BBC from recent staff cuts. Yet it remains confident in the belief that the BBC will maintain it, in spite of the vagaries of fashion, because the Corporation believes that such programmes deserve a place in the schedules of any broadcaster with pretensions of providing a Public Service. In due course, similar specialist Units were also established in London, in order to produce programmes on archaeology and history, on the arts, on music and on science. They too, at one time, had their successes. But they have not survived as well as the Unit in Bristol. The statutory requirement that a certain percentage of programmes must come from independent producers has reduced in-house production and the Units necessarily shrank proportionately in size. As they dwindled, so the critical mass of their production expertise has diminished. The continuity of their archives has been broken, they have lost the close touch they once had worldwide with their subjects and they are no longer regarded internationally as the centres of innovation and expertise that they once were.

When Birt gets up and says the whole of the BBC was a creative mess and it was wasteful, I never saw any evidence of that. I absolutely know it wasn’t so in my time. Producers now spend all their time worrying about money, and the thing has suffered for it.

In 2008, he criticised the BBC’s television schedules:

I have to say that there are moments when I wonder — moments when its two senior networks, first set up as a partnership, schedule simultaneously programmes of identical character, thereby contradicting the very reason that the BBC was given a second network. Then there are times when both BBC One and BBC Two, intoxicated by the sudden popularity of a programme genre, allow that genre to proliferate and run rampant through the schedules. The result is that other kinds of programmes are not placed, simply because of a lack of space. Do we really require so many gardening programmes, make-over programmes or celebrity chefs? Is it not a scandal in this day and age, that there seems to be no place for continuing series of programmes about science or serious music or thoughtful in-depth interviews with people other than politicians?

In 2009, Attenborough commented on the general state of British television, describing the newly introduced product placement on commercial television as something he considered an "appalling" idea 20 years earlier:

I think it's in great trouble. The whole system on which it was built — a limited number of networks, with adequate funding — is under threat. That funding is no longer there. As stations proliferate, so audiences are reduced. The struggle for audiences becomes ever greater, while money diminishes. I think that's a fair recipe for trouble. Inevitably, this has an impact on the BBC … Fortunately, the BBC doesn't think natural history programmes must compete with ''Strictly Come Dancing'' in terms of audience. The BBC says, 'Make proper, responsible natural history programmes.'

Other causes

In May 2005, Attenborough was appointed as patron of the UK's Blood Pressure Association, which provides information and support to people with hypertension.

Attenborough is also an honorary member of BSES Expeditions, a youth development charity that operates challenging scientific research expeditions to remote wilderness environments.

Parodies and artistic portrayals

Attenborough's accent and hushed, excited delivery have been the subject of frequent parodies by comedians, most notably Spike Milligan, Marty Feldman and ''The Goodies''. Attenborough is portrayed by Michael Palin in the final episode of ''Monty Python's Flying Circus'', where he searches the African jungle for the legendary Walking Tree of Dahomey (''Quercus Nicholas Parsonus''), sweating excessively and accompanied by native guides wearing saxophones. David Attenborough has also been parodied in a series of GEICO insurance commercials, showing a nature show host, clearly patterned on Attenborough.

"Time Flies", a sketch by David Ives, features a pair of anthropomorphic mayflies engaging in a courtship ritual, while watching themselves on television in a documentary narrated by David Attenborough.

The character of Nigel Thornberry, a nature documentarian on Nickelodeon's The Wild Thornberrys is strongly influenced by Attenborough.

He has also been parodied by the Australia 1980s sketch show The Comedy Company where Ian McFadyen portrays a character called David Rabbitborough.

The video game ''Discworld'', based on the series of books by Terry Pratchett, parodies his unique delivery to explain different aspects of the Discworld Universe, such as L-Space.

Mythbusters host Adam Savage often imitates Attenborough when speaking about his co-host Jamie Hyneman, which Jamie verified in the YouTube Special Episode.

Filmography

Attenborough is known foremost for writing and presenting the ten ''Life'' series, which are presented in chronological order below:

''Life on Earth'' (1979)

''The Living Planet'' (1984)

''The Trials of Life'' (1990)

''Life in the Freezer'' (1993)

''The Private Life of Plants'' (1995)

''The Life of Birds'' (1998)

''The Life of Mammals'' (2002)

''Life in the Undergrowth'' (2005)

''Life in Cold Blood'' (2008)

''First Life'' (2010)

His voice is synonymous with wildlife documentaries for British audiences, and the principal series with which his narration is associated include:

''Wildlife on One'' (1977–2005)

''BBC Wildlife Specials'' (1995–2008)

''The Blue Planet'' (2001)

''Planet Earth'' (British version) (2006)

''Nature's Great Events'' (2009)

''Life (2009)

''The Frozen Planet'' (2011) (in production)

Books

Bibliography

''Zoo Quest to Guyana'' (Lutterworth Press, 1956)

''Zoo Quest for a Dragon'' (Lutterworth Press, 1957)

* (book club edition with 85 extra pages, ''Quest for the Paradise Birds'', 1959)

''Zoo Quest in Paraguay'' (Lutterworth Press, 1959)

* paperback (Penguin Books, 1982)

''Quest in Paradise'' (1960)

''People of Paradise'' (Harper & Brothers, 1960)

''Zoo Quest to Madagascar'' (1961)

''Quest Under Capricorn'' (1963)

''Fabulous Animals'' (BBC, 1975) ISBN 0-563-17006-9

''The Tribal Eye'' (1976)

''Life on Earth'' (1979)

''Discovering Life on Earth'' (1981)

''Journeys to the past: Travels in New Guinea, Madagascar, and the northern territory of Australia'' (1983) Penguin Books ISBN 0-14-00.64133

''The Living Planet'' (1984)

''The First Eden: The Mediterranean World and Man'' (1987)(Little Brown & Co (T); 1st American ed edition (March 1990))

''The Atlas of the Living World'' (1989)

''The Trials of Life'' (Collins, 1990) ISBN 0-00-219912-2

''The Private Life of Plants'' (BBC Books, 1994) ISBN 0-563-37023-8

''The Life of Birds'' (BBC Books, 1998) ISBN 0-563-38792-0

''The Life of Mammals'' (BBC Books, 2002) ISBN 0-563-53423-0

''Life on Air: Memoirs of a Broadcaster'' (autobiography; 2002) ISBN 0-563-53461-3

* paperback: ISBN 0-563-48780-1

* updated edition: ISBN 9781846076527

''Life in the Undergrowth'' (BBC Books, 2005) ISBN 0-563-52208-9

''Amazing Rare Things - The Art of Natural History in the Age of Discovery'' with Susan Owens, Martin Clayton and Rea Alexandratos (The Royal Collection, 2007) Hardback - ISBN 978-1-902163-46-8; Softback - ISBN 978-1-902163-99-4

''Life in Cold Blood'' (BBC Books, 2007) ISBN 9780563539223

''Life Stories'', (HarperCollins Publishers, 2009) ISBN 9780007338832

''New Life Stories'', (Collins, 2011) ISBN 9780007425129

Introductions

Attenborough has written the introduction or foreword for a number of books, including:

''African Jigsaw: A Musical Entertainment'', Peter Rose and Anne Conlon (published: 1986, Weinberger)

''Tomorrow Is Too Late'', Various (The Macmillan Press, 1990)

''Life in the Freezer: Natural History of the Antarctic'', Alastair Fothergill (BBC Books, 1993), ISBN 0-563-36431-9

''Birds of Paradise: Paradisaeidae'' (Bird Families of the World series) Clifford B. Frith, Bruce M. Beehler, William T. Cooper (Illustrator) (Oxford University Press, 1998) ISBN 0-19-854853-2

''The Blue Planet'', Andrew Byatt, Alastair Fothergill, Martha Holmes (BBC Books, 2001) ISBN 0-563-38498-0.

''Light on the Earth'' (BBC Books, 2005), two decades of winning images from the Wildlife Photographer of the Year competition, ISBN 0-563-52260-7

''Planet Earth'', Alastair Fothergill (BBC Books, 2006), ISBN 0-563-52212-7

Audio recordings

''Tarka the Otter'' by Henry Williamson (available on audiocassette, 1978)

''Yanomamo'' (musical entertainment, 1983) by Peter Rose and Anne Conlon; on-stage narration and published audio recording

''Ocean World'' (musical entertainment, 1990) by Peter Rose and Anne Conlon; on-stage narration (including at The Royal Festival Hall), for audio recording and video broadcast (both published)

''Peter and the Wolf'' for BBC Music Magazine (free CD with the June 2000 issue).

In addition, Attenborough has recorded some of his own works in audiobook form, including ''Life on Earth'', ''Zoo Quest for a Dragon'' and his autobiography ''Life on Air: Memoirs of a Broadcaster''.

References

External links

BBC Books David Attenborough website

BBC ''Life on Air'' website

BBC biography

Wildfilmhistory.org biography

BBC interviews with Attenborough in 1976 and 1998

PBS interview with Attenborough in 1998

A Life in TV: Sir David Attenborough, BAFTA

BBC Wildlife Finder - David Attenborough's favourite moments

Tribute from the World Land Trust

Category:1926 births Category:Living people Category:Alumni of Clare College, Cambridge Category:Alumni of University College London Category:BBC Two controllers Category:Commanders of the Order of the British Empire Category:Commanders of the Royal Victorian Order Category:English agnostics Category:English conservationists Category:English environmentalists Category:English television personalities Category:Fellows of the American Academy of Arts and Sciences Category:Fellows of the Royal Society Category:Fellows of the Society of Antiquaries of London Category:Fellows of the Zoological Society of London Category:Kalinga Prize recipients Category:Knights Bachelor Category:Members of the Linnean Society of London Category:Members of the Order of the Companions of Honour Category:Members of the Order of Merit Category:People associated with the University of Leicester Category:People from Leicester Category:People from London Category:Presenters of the Royal Institution Christmas Lectures Category:British Book Award winners

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