The term "organism" (Greek ὀργανισμός – organismos, from Ancient Greek ὄργανον – organon "organ, instrument, tool") first appeared in the English language in 1701 and took on its current definition by 1834 (Oxford English Dictionary). It is directly related to the term "organization". There is a long tradition of defining organisms as self-organizing beings.
There has been a great deal of recent controversy about the best way to define the organism and indeed about whether or not such a definition is necessary. Several contributions are responses to the suggestion that the category of "organism" may well not be adequate in biology.
Scientific classification in biology considers organisms synonymous with "life on Earth". Based on cell type, organisms may be divided into the prokaryotic and eukaryotic groups. The prokaryotes represent two separate domains, the Bacteria and Archaea. Eukaryotic organisms, with a membrane-bounded cell nucleus, also contain organelles, namely mitochondria and (in plants) plastids, generally considered to be derived from endosymbiotic bacteria. Fungi, animals and plants are examples of species that are eukaryotes.
More recently a clade, Neomura, has been proposed, which groups together the Archaea and Eukarya. Neomura is thought to have evolved from Bacteria, more specifically from Actinobacteria.
Chambers Online Reference provides a broad definition: "any living structure, such as a plant, animal, fungus or bacterium, capable of growth and reproduction".
In multicellular terms, "organism" usually describes the whole hierarchical assemblage of systems (for example circulatory, digestive, or reproductive) themselves collections of organs; these are, in turn, collections of tissues, which are themselves made of cells. In some plants and the nematode Caenorhabditis elegans, individual cells are totipotent.
A superorganism is an organism consisting of many individuals working together as a single functional or social unit.
Taxa are ranked groups of organisms, which run from the general (domain) to the specific (species). A broad scheme of ranks in hierarchical order is:
To give an example, Homo sapiens is the Latin binomial equating to modern humans. All members of the species sapiens are, at least in theory, genetically able to interbreed. Several species may belong to a genus, but the members of different species within a genus are unable to interbreed to produce fertile offspring. Homo, however, only has one surviving species (sapiens), Homo erectus, Homo neanderthalensis, etc. having become extinct thousands of years ago. Several genera belong to the same family and so on up the hierarchy. Eventually, the relevant kingdom (Animalia, in the case of humans) is placed into one of the three domains depending upon certain genetic and structural characteristics.
All living organisms known to science are given classification by this system such that the species within a particular family are more closely related and genetically similar than the species within a particular phylum.
Organisms clearly owe their origin, metabolism, and many other internal functions to chemical phenomena, especially the chemistry of large organic molecules. Organisms are complex systems of chemical compounds that, through interaction and environment, play a wide variety of roles.
Organisms are semi-closed chemical systems. Although they are individual units of life (as the definition requires) they are not closed to the environment around them. To operate they constantly take in and release energy. Autotrophs produce usable energy (in the form of organic compounds) using light from the sun or inorganic compounds while heterotrophs take in organic compounds from the environment.
The primary chemical element in these compounds is carbon. The physical properties of this element such as its great affinity for bonding with other small atoms, including other carbon atoms, and its small size making it capable of forming multiple bonds, make it ideal as the basis of organic life. It is able to form small three-atom compounds (such as carbon dioxide), as well as large chains of many thousands of atoms that can store data (nucleic acids), hold cells together, and transmit information (protein).
These protein functions have been recognized: # Enzymes, which catalyze all of the reactions of metabolism # Structural proteins, such as tubulin, or collagen # Regulatory proteins, such as transcription factors or cyclins that regulate the cell cycle # Signaling molecules or their receptors such as some hormones and their receptors # Defensive proteins, which can include everything from antibodies of the immune system, to toxins (e.g., dendrotoxins of snakes), to proteins that include unusual amino acids like canavanine
Lipids make up the membrane of cells that constitutes a barrier, containing everything within the cell and preventing compounds from freely passing into, and out of, the cell. In some multicellular organisms they serve to store energy and mediate communication between cells. Carbohydrates also store and transport energy in some organisms, but are more easily broken down than lipids.
There are two types of cells, eukaryotic and prokaryotic. Prokaryotic cells are usually singletons, while eukaryotic cells are usually found in multicellular organisms. Prokaryotic cells lack a nuclear membrane so DNA is unbound within the cell, eukaryotic cells have nuclear membranes.
All cells, whether prokaryotic or eukaryotic, have a membrane, which envelops the cell, separates its interior from its environment, regulates what moves in and out, and maintains the electric potential of the cell. Inside the membrane, a salty cytoplasm takes up most of the cell volume. All cells possess DNA, the hereditary material of genes, and RNA, containing the information necessary to build various proteins such as enzymes, the cell's primary machinery. There are also other kinds of biomolecules in cells.
All cells share several abilities:
Today, there is debate over whether or not all organisms descended from a common ancestor, or a "last universal ancestor" (LUA), also called the "last universal common ancestor" (LUCA). The universality of genetic coding suggests common ancestry. For example, every living cell makes use of nucleic acids as its genetic material, and uses the same twenty amino acids as the building blocks for proteins, although exceptions to the basic twenty amino acids have been found. However, throughout history groupings based on appearance or function of species have sometimes been polyphyletic due to convergent evolution.
The "last universal ancestor" (LUA), or "last universal common ancestor" (LUCA), is the name given to the hypothetical unicellular organism or single cell that gave rise to all life on Earth 3.5 to 3.8 billion years ago; however, this hypothesis has since been refuted on many grounds. For example, it was once thought that the genetic code was universal (see: Universal genetic code), but many variations have been discovered including various alternative mitochondrial codes. Back in the early 1970s, evolutionary biologists thought that a given piece of DNA specified the same protein subunit in every living thing, and that the genetic code was thus universal. This was interpreted as evidence that every organism had inherited its genetic code from a single common ancestor, aka, an LUCA. In 1979, however, exceptions to the code were found in mitochondria, the tiny energy factories inside cells. Researchers studying human mitochondrial genes discovered that they used an alternative code, and many slight variants have been discovered since, In certain proteins, non-standard amino acids are substituted for standard stop codons, depending upon associated signal sequences in the messenger RNA: UGA can code for selenocysteine and UAG can code for pyrrolysine. Selenocysteine is now viewed as the 21st amino acid, and pyrrolysine is viewed as the 22nd. A detailed description of variations in the genetic code can be found at the NCBI web site.
Information about the early development of life includes input from many different fields, including geology and planetary science. These sciences provide information about the history of the Earth and the changes produced by life. However, a great deal of information about the early Earth has been destroyed by geological processes over the course of time.
stromatolites in the Siyeh Formation, Glacier National Park. In 2002, William Schopf of UCLA published a controversial paper in the journal Nature arguing that formations such as this possess 3.5 billion year old fossilized algae microbes. If true, they would be the earliest known life on earth.]] Not much is known about the earliest developments in life. However, all existing organisms share certain traits, including cellular structure and genetic code. Most scientists interpret this to mean all existing organisms share a common ancestor, which had already developed the most fundamental cellular processes, but there is no scientific consensus on the relationship of the three domains of life (Archaea, Bacteria, Eukaryota) or the origin of life. Attempts to shed light on the earliest history of life generally focus on the behavior of macromolecules, particularly RNA, and the behavior of complex systems.
The emergence of oxygenic photosynthesis (around 3 billion years ago) and the subsequent emergence of an oxygen-rich, non-reducing atmosphere can be traced through the formation of banded iron deposits, and later red beds of iron oxides. This was a necessary prerequisite for the development of aerobic cellular respiration, believed to have emerged around 2 billion years ago.
In the last billion years, simple multicellular plants and animals began to appear in the oceans. Soon after the emergence of the first animals, the Cambrian explosion (a period of unrivaled and remarkable, but brief, organismal diversity documented in the fossils found at the Burgess Shale) saw the creation of all the major body plans, or phyla, of modern animals. This event is now believed to have been triggered by the development of the Hox genes. About 500 million years ago, plants and fungi colonized the land, and were soon followed by arthropods and other animals, leading to the development of today's land ecosystems.
The evolutionary process may be exceedingly slow. Fossil evidence indicates that the diversity and complexity of modern life has developed over much of the history of the earth. Geological evidence indicates that the Earth is approximately 4.6 billion years old. Studies on guppies by David Reznick at the University of California, Riverside, however, have shown that the rate of evolution through natural selection can proceed 10 thousand to 10 million times faster than what is indicated in the fossil record. Such comparative studies however are invariably biased by disparities in the time scales over which evolutionary change is measured in the laboratory, field experiments, and the fossil record.
Sequence comparisons suggest recent horizontal transfer of many genes among diverse species including across the boundaries of phylogenetic "domains". Thus determining the phylogenetic history of a species can not be done conclusively by determining evolutionary trees for single genes.
Biologist Gogarten suggests "the original metaphor of a tree no longer fits the data from recent genome research", therefore "biologists [should] use the metaphor of a mosaic to describe the different histories combined in individual genomes and use [the] metaphor of a net to visualize the rich exchange and cooperative effects of HGT among microbes."
The J. Craig Venter Institute has recently assembled a synthetic bacterial genome, Mycoplasma genitalium, by using recombination in yeast of 25 overlapping DNA fragments in a single step. "The use of yeast recombination greatly simplifies the assembly of large DNA molecules from both synthetic and natural fragments." Other companies, such as Synthetic Genomics, have already been formed to take advantage of the many commercial uses of custom designed genomes.
This text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
Name | Noah |
---|---|
Birth date | approx. 2900 BC |
Death date | approx. 2000 BC |
Venerated in | JudaismChristianityIslamMandaeismBaha'i Faith |
Caption | Noah's sacrifice by Daniel Maclise |
Birth place | Mesopotamia (?) |
Titles | Antediluvian Patriarch, Prophet, Holy Forefather, Constructor of the Ark, 'Grateful Servant of God', 'Preacher of Righteousness' |
Influenced | Many Jews, Christians and Muslims |
:The following section is a summary of the Book of Genesis, chapters 6–9. Noah was the son of Lamech who named him Noah, saying, "This same shall comfort us in our work and in the toil of our hands, which cometh from the ground which the LORD hath cursed." In his five hundredth year Noah had three sons, Shem, Ham, and Japheth. In his six hundredth year God, saddened at the wickedness of mankind, sent a great deluge to destroy all life, but instructed Noah, a man "righteous in his generation," to build an ark and save a remnant of life from the Flood.
After the Flood, "Noah was the first tiller of the soil. He planted a vineyard; and he drank of the wine." Noah's son Ham saw his father naked in his father's tent, and told his brothers, and so Noah cursed Ham's son Canaan, giving his land to Shem.
Noah died 350 years after the Flood, at the age of 950, the last of the immensely long-lived antediluvian Patriarchs. The maximum human lifespan, as depicted by the Bible, diminishes rapidly thereafter, from as much as 900 years to the 120 years of Moses.
The righteousness of Noah is the subject of much discussion among the rabbis. The description of Noah as "righteous in his generation" implied to some that his perfection was only relative: In his generation of wicked people, he could be considered righteous, but in the generation of a tzadik like Abraham, he would not be considered so righteous. They point out that Noah did not pray to God on behalf of those about to be destroyed, as Abraham prayed for the wicked of Sodom and Gomorrah. In fact, Noah is never seen to speak; he simply listens to God and acts on his orders. This led such commentators to offer the figure of Noah as "the man in a fur coat," who ensured his own comfort while ignoring his neighbour. Others, such as the medieval commentator Rashi, held on the contrary that the building of the Ark was stretched over 120 years, deliberately in order to give sinners time to repent. Rashi interprets his father's statement of the naming of Noah (in Hebrew נֹחַ) “This one will comfort (in Hebrew– yeNaHamainu יְנַחֲמֵנו) from our work and our hands sore from the land that the Lord had cursed”, by saying Noah heralded a new era of prosperity, when there was easing (in Hebrew – nahah - נחה) from the curse from the time of Adam when the Earth produced thorns and thistles even where men sowed wheat and that Noah then introduced the plow.
Noah's three sons were generally interpreted in medieval Christianity as the founders of the populations of the three known continents, Japheth/Europe, Shem/Asia, and Ham/Africa, although a rarer variation held that they represented the three classes of medieval society - the priests (Shem), the warriors (Japheth), and the peasants (Ham). In the 18th and 19th centuries the view that Ham's sons in general had been literally "blackened" by the curse of Noah was cited as justification for black slavery.
The Qur'an focuses on several instances from Noah's life more than others, and one of the most significant events is the Deluge. God makes a covenant with Noah just as with Abraham, Moses, Jesus and Muhammad later on (XXXIII: 7). Noah is later reviled by his people and reproached by them for being a mere human messenger and not an angel (X: 72-74). Moreover, the people of Noah mock Noah's words and call him a liar (VII: 62) and even suggest that Noah is possessed by a devil when the prophet ceases to preach (LIV: 9). Only the lowest in the community join Noah in believing in God's message (XI: 29), and Noah's narrative further describes him preaching both in private and public. The Qur'an narrates that Noah received a revelation to build an Ark, after his people refused to believe in his message and hear the warning. The narrative goes onto describe that waters poured forth from the Heavens, destroying all the sinners, including one of Noah's own sons, who refused to believe his father. After the Great Flood ceased, the Ark rested atop Mount Judi (XI: 27-51).
More broadly, Genesis seems to contain two accounts concerning Noah, the first making him the hero of the Flood, the second representing him as a husbandman who planted a vineyard. This has led some scholars to believe that Noah was believed by the ancients to be the inventor of wine, in keeping with the statement at that Lamech "called his name Noah, saying, 'Out of the ground which the Lord has cursed this one shall bring us relief from our work and from the toil of our hands.'"
.]] Noah's great grandfather Enoch, according to , at the end of his 365 years, "walked with God, and was not, for God took him." The prevalent Hebrew understanding of the text meaning that he was taken to heaven to be with God. Of Enoch, the only of the ten pre-Flood Patriarchs not reported to have died, it is not explicitly stated where he is taken in Genesis. However, states that "by faith Enoch was taken from this life, so that he did not experience death; he could not be found, because God had taken him away. For before he was taken, he was commended as one who pleased God." Enoch and Elijah (in : "the LORD was about to take Elijah up to heaven in a whirlwind") are two people described as experiencing assumption (not seeing death). In a late Apocryphal tradition, Methuselah is reported to have visited Enoch at the end of the Earth, where he dwelt with the angels, immortal. The details bring to mind Utnapishtim, a figure from the Mesopotamian Epic of Gilgamesh - the hero Gilgamesh, after long and arduous travel, finds Utnapishtim living in the paradise of Dilmun / Bahrain at the end of the Earth, where he has been granted eternal life by the gods. (Gilgamesh's reason for seeking out Utnapishtim, incidentally, is to learn the secret of immortality - like Methuselah, he comes close to the gift but fails to achieve it). Utnapishtim then tells how he survived a great flood, and how he was afterwards granted immortality by the gods. It has been suggested that the Flood story may originally have belonged to Enoch.
Lamech's statement that Noah will be named "rest" because "out of the ground which the Lord has cursed this one shall bring us relief from our work and from the toil of our hands," has another faint parallel in Babylonian mythology: the gods grew tired of working, digging the channels of the rivers, and so the god Enki created man from clay and blood and spit to do the work for them. Enki fell in love with his creation, and later warned Utnapishtim that the other gods planned to send a flood to destroy all life, and advised him on how to construct his ark.
Noah is also often compared to Deucalion, the son of Prometheus and Pronoia in Greek mythology. Like Noah, Deucalion is a wine maker or wine seller; he is forewarned of the flood (this time by Zeus); he builds an ark and staffs it with creatures - and when he completes his voyage, gives thanks and takes advice from the gods on how to repopulate the Earth. Deucalion also sends a pigeon to find out about the situation of the world and the bird return with an olive branch. This and some other examples of apparent comparison between Greek myths and the "key characters" in the Old Testament/Torah have led recent Biblical scholars to suggest a Hellenistic influence in the composition of the earlier portions of the Hebrew Bible.
Category:Abrahamic mythology Category:Legendary progenitors Category:Longevity traditions Category:Noahides Category:Old Testament saints Category:People celebrated in the Lutheran liturgical calendar Category:Torah people Category:Noah's Ark *
This text is licensed under the Creative Commons CC-BY-SA License. This text was originally published on Wikipedia and was developed by the Wikipedia community.
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