Biodiversity is the degree of variation of life forms within a given ecosystem, biome, or an entire planet. Biodiversity is a measure of the health of ecosystems. Greater biodiversity implies greater health. Biodiversity is in part a function of climate. In terrestrial habitats, tropical regions are typically rich whereas polar regions support fewer species.
Rapid environmental changes typically cause extinctions. One estimate is that less than 1% of the species that have existed on Earth are extant.
Since life began on Earth, five major mass extinctions and several minor events have led to large and sudden drops in biodiversity. The Phanerozoic eon (the last 540 million years) marked a rapid growth in biodiversity via the Cambrian explosion—a period during which nearly every phylum of multicellular organisms first appeared. The next 400 million years included repeated, massive biodiversity losses classified as mass extinction events. In the Carboniferous, rainforest collapse led to a great loss of plant and animal life. The Permian–Triassic extinction event, 251 million years ago, was the worst; vertebrate recovery took 30 million years. The most recent, the Cretaceous–Tertiary extinction event, occurred 65 million years ago, and has often attracted more attention than others because it resulted in the extinction of the dinosaurs.
The period since the emergence of humans has displayed an ongoing biodiversity reduction and an accompanying loss of genetic diversity. Named the Holocene extinction, the reduction is caused primarily by human impacts, particularly habitat destruction. Biodiversity's impact on human health is a major international issue.
The United Nations designated 2011-2020 as the United Nations Decade on Biodiversity.
The term's contracted form biodiversity may have been coined by W.G. Rosen in 1985 while planning the 1986 National Forum on Biological Diversity organized by the National Research Council (NRC). It first appeared in a publication in 1988 when entomologist E. O. Wilson used it as the title of the proceedings of that forum.
Since this period the term has achieved widespread use among biologists, environmentalists, political leaders, and concerned citizens.
A similar term in the United States is "natural heritage." It predates the others and is more accepted by the wider audience interested in conservation. Broader than biodiversity, it includes geology and landforms (geodiversity).
"Biological diversity" or "biodiversity" can have many interpretations. It is most commonly used to replace the more clearly defined and long established terms, species diversity and species richness. Biologists most often define biodiversity as the "totality of genes, species, and ecosystems of a region". An advantage of this definition is that it seems to describe most circumstances and presents a unified view of the traditional three levels at which biological variety has been identified:
This multilevel construct is consistent with Dasmann and Lovejoy. An explicit definition consistent with this interpretation was first given in a paper by Bruce A. Wilcox commissioned by the International Union for the Conservation of Nature and Natural Resources (IUCN) for the 1982 World National Parks Conference. Wilcox's definition was "Biological diversity is the variety of life forms...at all levels of biological systems (i.e., molecular, organismic, population, species and ecosystem)..." The 1992 United Nations Earth Summit defined "biological diversity" as "the variability among living organisms from all sources, including, 'inter alia', terrestrial, marine, and other aquatic ecosystems, and the ecological complexes of which they are part: this includes diversity within species, between species and of ecosystems". This definition is used in the United Nations Convention on Biological Diversity.
One textbook's definition is "variation of life at all levels of biological organization".
Geneticists define it as the diversity of genes and organisms. They study processes such as mutations, gene transfer, and genome dynamics that generate evolution.
Selection bias amongst researchers may contribute to biased empirical research for modern estimates of biodiversity. In 1768 Rev. Gilbert White succinctly observed of his Selborne, Hampshire "all nature is so full, that that district produces the most variety which is the most examined."
Biodiversity is not evenly distributed. Flora and fauna diversity depends on climate, altitude, soils and the presence of other species. Diversity consistently measures higher in the tropics and in other localized regions such as Cape Floristic Province and lower in polar regions generally. In 2006 many species were formally classified as rare or endangered or threatened; moreover, scientists have estimated that millions more species are at risk which have not been formally recognized. About 40 percent of the 40,177 species assessed using the IUCN Red List criteria are now listed as threatened with extinction—a total of 16,119.
Even though terrestrial biodiversity declines from the equator to the poles, this characteristic is unverified in aquatic ecosystems, especially in marine ecosystems. In addition, several assessments reveal tremendous diversity in higher latitudes. Generally terrestrial biodiversity is up to 25 times greater than ocean biodiversity.
A biodiversity hotspot is a region with a high level of endemic species. Hotspots were first named in 1988 by Dr. Norman Myers. Many hotspots have large nearby human populations. Most hotspots are located in the tropics and most of them are forests.
Brazil's Atlantic Forest is considered one such hotspot, containing roughly 20,000 plant species, 1,350 vertebrates, and millions of insects, about half of which occur nowhere else. The island of Madagascar, particularly the unique Madagascar dry deciduous forests and lowland rainforests, possess a high ratio of endemism. Since the island separated from mainland Africa 65 million years ago, many species and ecosystems have evolved independently. Indonesia's 17,000 islands cover contain 10% of the world's flowering plants, 12% of mammals and 17% of reptiles, amphibians and birds—along with nearly 240 million people. Many regions of high biodiversity and/or endemism arise from specialized habitats which require unusual adaptations, for example alpine environments in high mountains, or Northern European peat bogs.
Biodiversity is the result of 3.5 billion years of evolution. The origin of life has not been definitely established by science, however some evidence suggests that life may already have been well-established only a few hundred million years after the formation of the Earth. Until approximately 600 million years ago, all life consisted of archaea, bacteria, protozoans and similar single-celled organisms.
The history of biodiversity during the Phanerozoic (the last 540 million years), starts with rapid growth during the Cambrian explosion—a period during which nearly every phylum of multicellular organisms first appeared. Over the next 400 million years or so, invertebrate diversity showed little overall trend, and vertebrate diversity shows an overall exponential trend., whereas others consider the fossil record reasonably reflective of the diversification of life. Diversity appears to increase continually in the absence of natural selection.
Most biologists agree however that the period since human emergence is part of a new mass extinction, named the Holocene extinction event, caused primarily by the impact humans are having on the environment. It has been argued that the present rate of extinction is sufficient to eliminate most species on the planet Earth within 100 years.
New species are regularly discovered (on average between 5–10,000 new species each year, most of them insects) and many, though discovered, are not yet classified (estimates are that nearly 90% of all arthropods are not yet classified). climate (e.g., sequestration), water purification, pollination, and prevention of erosion.
Non-material benefits include spiritual and aesthetic values, knowledge systems and the value of education.
The reservoir of genetic traits present in wild varieties and traditionally grown landraces is extremely important in improving crop performance. Important crops, such as potato, banana and coffee, are often derived from only a few genetic strains. Improvements in crop species over the last 250 years have been largely due to incorporating genes from wild varieties and species into cultivars. Crop breeding for beneficial traits has helped to more than double crop production in the last 50 years as a result of the Green Revolution. A biodiverse environment preserves the genome from which such productive genes are drawn.
Crop diversity aids recovery when the dominant cultivar is attacked by a disease or predator:
Monoculture was a contributing factor to several agricultural disasters, including the European wine industry collapse in the late 19th century, and the US Southern Corn Leaf Blight epidemic of 1970.
Although about 80 percent of humans' food supply comes from just 20 kinds of plants, humans use at least 40,000 species. Many people depend on these species for food, shelter, and clothing. Earth's surviving biodiversity provides resources for increasing the range of food and other products suitable for human use, although the present extinction rate shrinks that potential. This issue is closely linked with the issue of climate change, as many of the anticipated health risks of climate change are associated with changes in biodiversity (e.g. changes in populations and distribution of disease vectors, scarcity of fresh water, impacts on agricultural biodiversity and food resources etc.) Some of the health issues influenced by biodiversity include dietary health and nutrition security, infectious disease, medical science and medicinal resources, social and psychological health. Biodiversity is also known to have an important role in reducing disaster risk, and in post-disaster relief and recovery efforts.
Biodiversity provides critical support for drug discovery and the availability of medicinal resources. A significant proportion of drugs are derived, directly or indirectly, from biological sources: at least 50% of the pharmaceutical compounds on the US market are derived from plants, animals, and micro-organisms, while about 80% of the world population depends on medicines from nature (used in either modern or traditional medical practice) for primary healthcare. Marine ecosystems are particularly important, although inappropriate bioprospecting can increase biodiversity loss, as well as violating the laws of the communities and states from which the resources are taken. Higher biodiversity also limits the spread of infectious diseases as many different species act as buffers to them.
Popular activities such as gardening, fishkeeping and specimen collecting strongly depend on biodiversity. The number of species involved in such pursuits is in the tens of thousands, though the majority do not enter commerce.
The relationships between the original natural areas of these often exotic animals and plants and commercial collectors, suppliers, breeders, propagators and those who promote their understanding and enjoyment are complex and poorly understood. The general public responds well to exposure to rare and unusual organisms, reflecting their inherent value.
Philosophically it could be argued that biodiversity has intrinsic aesthetic and spiritual value to mankind in and of itself. This idea can be used as a counterweight to the notion that tropical forests and other ecological realms are only worthy of conservation because of the services they provide.
Biodiversity supports many ecosystem services that are often not readily visible. It plays a part in regulating the chemistry of our atmosphere and water supply. Biodiversity is directly involved in water purification, recycling nutrients and providing fertile soils. Experiments with controlled environments have shown that humans cannot easily build ecosystems to support human needs; for example insect pollination cannot be mimicked, and that activity alone represents tens of billions of dollars in ecosystem services per year to humankind.
Ecosystem stability is also positively related to biodiversity, protecting against disruption by extreme weather or human exploitation.
s on the sea ice of the Arctic Ocean, near the North Pole.]]
Since the rate of extinction has increased, many extant species may become extinct before they are described.
Habitat destruction has played a key role in extinctions, especially related to tropical forest destruction. Factors contributing to habitat loss are: overpopulation, deforestation, pollution (air pollution, water pollution, soil contamination) and global warming or climate change.
Habitat size and numbers of species are systematically related. Physically larger species and those living at lower latitudes or in forests or oceans are more sensitive to reduction in habitat area. Conversion to "trivial" standardized ecosystems (e.g., monoculture following deforestation) effectively destroys habitat for the more diverse species that preceded the conversion. In some countries lack of property rights or lax law/regulatory enforcement necessarily leads to biodiversity loss (degradation costs having to be supported by the community).
A 2007 study conducted by the National Science Foundation found that biodiversity and genetic diversity are codependent—that diversity among species requires diversity within a species, and vice versa. "If any one type is removed from the system, the cycle can break down, and the community becomes dominated by a single species." At present, the most threatened ecosystems are found in fresh water, according to the Millennium Ecosystem Assessment 2005, which was confirmed by the "Freshwater Animal Diversity Assessment", organised by the biodiversity platform, and the French Institut de recherche pour le développement (MNHNP).
Co-extinctions are a form of habitat destruction. Co-extinction occurs when the extinction or decline in one accompanies the other, such as in plants and beetles.
Not all introduced species are invasive, nor all invasive species deliberately introduced. In cases such as the zebra mussel, invasion of US waterways was unintentional. In other cases, such as mongooses in Hawaii, the introduction is deliberate but ineffective (nocturnal rats were not vulnerable to the diurnal mongoose). In other cases, such as oil palms in Indonesia and Malaysia, the introduction produces substantial economic benefits, but the benefits are accompanied by costly unintended consequences.
Finally, an introduced species may unintentionally injure a species that depends on the species it replaces. In Belgium, Prunus spinosa from Eastern Europe leafs much sooner than its West European counterparts, disrupting the feeding habits of the Thecla betulae butterfly (which feeds on the leaves). Introducing new species often leaves endemic and other local species unable to compete with the exotic species and unable to survive. The exotic organisms may be predators, parasites, or may simply outcompete indigenous species for nutrients, water and light.
At present, several countries have already imported so many exotic species, particularly agricultural and ornamental plants, that the own indigenous fauna/flora may be outnumbered.
Endemic species can be threatened with extinction through the process of genetic pollution, i.e. uncontrolled hybridization, introgression and genetic swamping. Genetic pollution leads to homogenization or replacement of local genomes as a result of either a numerical and/or fitness advantage of an introduced species. Hybridization and introgression are side-effects of introduction and invasion. These phenomena can be especially detrimental to rare species that come into contact with more abundant ones. The abundant species can interbreed with the rare species, swamping its gene pool. This problem is not always apparent from morphological (outward appearance) observations alone. Some degree of gene flowis normal adaptation, and not all gene and genotype constellations can be preserved. However, hybridization with or without introgression may, nevertheless, threaten a rare species' existence.
Overexploitation occurs when a resource is consumed at an unsustainable rate. This occurs on land in the form of overhunting, excessive logging, poor soil conservation in agriculture and the illegal wildlife trade. Joe Walston, director of the Wildlife Conservation Society’s Asian programs, called the latter the "single largest threat" to biodiversity in Asia. The international trade of endangered species is second in size only to drug trafficking.
About 25% of world fisheries are now overfished to the point where their current biomass is less than the level that maximizes their sustainable yield.
The overkill hypothesis explains why earlier megafaunal extinctions occurred within a relatively short period of time. This can be connected with human migration.
In agriculture and animal husbandry, the Green Revolution popularized the use of conventional hybridization to increase yield. Often hybridized breeds originated in developed countries and were further hybridized with local varieties in the developing world to create high yield strains resistant to local climate and diseases. Local governments and industry have been pushing hybridization. Formerly huge gene pools of various wild and indigenous breeds have collapsed causing widespread genetic erosion and genetic pollution. This has resulted in loss of genetic diversity and biodiversity as a whole.
(GM organisms) have genetic material altered by genetic engineering procedures such as recombinant DNA technology. GM crops have become a common source for genetic pollution, not only of wild varieties but also of domesticated varieties derived from classical hybridization.
Genetic erosion coupled with genetic pollution may be destroying unique genotypes, thereby creating a hidden crisis which could result in a severe threat to our food security. Diverse genetic material could cease to exist which would impact our ability to further hybridize food crops and livestock against more resistant diseases and climatic changes.
In 2004, an international collaborative study on four continents estimated that 10 percent of species would become extinct by 2050 because of global warming. "We need to limit climate change or we wind up with a lot of species in trouble, possibly extinct," said Dr. Lee Hannah, a co-author of the paper and chief climate change biologist at the Center for Applied Biodiversity Science at Conservation International.
of Aletsch Glacier in the Swiss Alps (situation in 1979, 1991 and 2002), due to global warming.]]
Conservation biology matured in the mid-20th century as ecologists, naturalists, and other scientists began to research and address issues pertaining to global biodiversity declines.
The conservation ethic advocates management of natural resources for the purpose of sustaining biodiversity in species, ecosystems, the evolutionary process, and human culture and society.
Conservation biology is reforming around strategic plans to protect biodiversity. Preserving global biodiversity is a priority in strategic conservation plans that are designed to engage public policy and concerns affecting local, regional and global scales of communities, ecosystems, and cultures. Action plans identify ways of sustaining human well-being, employing natural capital, market capital, and ecosystem services.
Exotic species removal allows less competitive species to recover their ecological niches. Exotic species that have become a pest can be identified taxonomically (e.g. with Digital Automated Identification SYstem (DAISY), using the barcode of life. Removal is practical only given large groups of individuals due to the econimic cost.
Once the preservation of the remaining native species in an area is assured. "missing" species can be identified and reintroduced using databases such as the Encyclopedia of Life and the Global Biodiversity Information Facility.
Other techniques include:
A second strategy focuses on areas that retain most of their original diversity, which typically require little or no restoration. These are typically non-urbanized, non-agricultural areas. Tropical areas often fit both criteria, given their natively high diversity and relative lack of development.
Biodiversity is taken into account in some political and judicial decisions: The relationship between law and ecosystems is very ancient and has consequences for biodiversity. It is related to private and public property rights. It can define protection for threatened ecosystems, but also some rights and duties (for example, fishing and hunting rights).
Global agreements such as the Convention on Biological Diversity), give "sovereign national rights over biological resources" (not property). The agreements commit countries to "conserve biodiversity", "develop resources for sustainability" and "share the benefits" resulting from their use. Biodiverse countries that allow bioprospecting or collection of natural products, expect a share of the benefits rather than allowing the individual or institution that discovers/exploits the resource to capture them privately. Bioprospecting can become a type of biopiracy when such principles are not respected.
Sovereignty principles can rely upon what is better known as Access and Benefit Sharing Agreements (ABAs). The Convention on Biodiversity implies informed consent between the source country and the collector, to establish which resource will be used and for what, and to settle on a fair agreement on benefit sharing.
Uniform approval for use of biodiversity as a legal standard has not been achieved, however. Bosselman argues that biodiversity should not be used as a legal standard, claiming that the remaining areas of scientific uncertainty cause unacceptable administrative waste and increase litigation without promoting preservation goals.
Category:Biology terminology Category:Environmental science Category:Habitat (ecology) terminology Category:Millennium Development Goals
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