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The penis (plural penises, penes) is a biological feature of male animals including both vertebrates (creatures with a backbone) and invertebrates. It is a reproductive, intromittent organ that additionally serves as the urinal duct in placental mammals.
As with nearly any aspect of the body involved in sexual or excretory functions, the penis is the subject of taboos, and there are many slang words and euphemisms for it, a particularly common and longstanding one being "cock".
The Latin word "phallus" (from Greek φαλλος) is sometimes used to describe the penis, although "phallus" originally was used to describe images, pictorial or carved, of the penis.
Pizzle, an archaic English word for penis, of Low German or Dutch origin, is is now used to denote the penis of a non human animal.
The adjectival form of the word penis is penile. This adjective is commonly used in describing various accessory structures of male copulatory organs found in many kinds of invertebrate animals.
A bone called the baculum or os penis is present in most mammals but absent in humans and horses.
The internal structures of the penis consist mainly of cavernous (erectile) tissue, which is a collection of blood sinusoids separated by sheets of connective tissue (trabeculae). Some animals have a lot of erectile tissue relative to connective tissue, for example horses. Because of this a horse's penis can enlarge more than a bull's penis. The urethra is on the ventral side of the body of the penis.
Stallions have a vascular penis. When non-erect, it is quite flaccid and contained within the prepuce (sheath). The retractor penis muscle is relatively underdeveloped. Erection and protrusion take place gradually, by the increasing tumescence of the erectile vascular tissue in the corpus cavernosum penis.
A bull has a fibro-elastic penis. It has a small diameter and is quite rigid when non-erect. There is a relatively small amount of erectile tissue and there is barely any enlargement after erection but the penis becomes more rigid. Protrusion is not affected much by erection, but more by relaxation of the retractor penis muscle and straightening out of the sigmoid flexure.
In the realm of absolute size, the smallest vertebrate penis belongs to the Common Shrew (5 mm or 0.2 inches). Accurate measurements of the blue whale are difficult to take because the whale's erect length can only be observed during mating.
Most marsupials, except for the two largest species of kangaroos, have a bifurcated penis. That is, it separates into two columns, and so the penis has two ends corresponding to the females' two vaginas. Neither marsupials nor monotremes possess a baculum.
Echidnas have a four-headed penis, but only two of the heads are used during mating. The other two heads "shut down" and do not grow in size. The heads used are swapped each time the mammal has sex.
Male specimens of the reptile order Squamata have two paired organs called hemipenes.
In some fishes, the gonopodium, andropodium, and claspers are intromittent organs (to introduce sperm into the female) developed from modified fins.
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In male insects, the structure analogous to a penis is known as aedeagus. The male copulatory organ of various lower invertebrate animals is often called the cirrus.
A number of invertebrate species have independently evolved the mating technique of traumatic insemination where the penis penetrates the female's abdomen and deposits sperm in the wound it produces. This has been most fully studied in bedbugs.
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